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Acta zoológica mexicana

 ISSN 2448-8445 ISSN 0065-1737

        03--2025

https://doi.org/10.21829/azm.2025.4112775 

Artículos originales

The velvet ants (Hymenoptera, Mutillidae) of Mexico: a taxonomic account

Los Mutílidos (Hymenoptera, Mutillidae) de México: una perspectiva taxonómica

Fernando Varela-Hernández1 
http://orcid.org/0000-0001-9945-4667

1Laboratorio de Sistemática Molecular, Escuela de Estudios Superiores del Jicarero, Universidad Autónoma del Estado de Morelos, C. P. 62909, Jojutla, Morelos, Mexico.

Abstract.

Digitalized data from different collections and information from the literature were analyzed to document the diversity of velvet ants of Mexico. A checklist of 369 species was compiled. This work has contributed to increasing the known species of Mexico by 40.30%. The most species-rich genus is Dasymutilla with 114 species and the less species-rich genera are Hoplognathoca, Myrmosula and Pertyella with one species each. The states with the most genera are Sonora, Chihuahua, and Coahuila and those with the most species are Sonora, Baja California, and Chihuahua. This study represents the first attempt to integrate information on the velvet ants occurring in Mexico and their geographic distribution per state.

Key words: Checklist; diversity; geographic distribution; Mexico; species

Resumen.

Datos digitalizados de diferentes colecciones y la información proveniente de literatura fue analizada para documentar la diversidad de mutílidos de México. Se generó una lista anotada de 369 especies. Este trabajo ha contribuido a incrementar la riqueza de especies conocida para México en un 40.30%. El género con mayor número de especies es Dasymutilla con 114 y los géneros menos diversos son Hoplognathoca, Myrmosula y Pertyella con una especie cada uno. Los estados con el mayor número de géneros son Sonora, Chihuahua y Coahuila y aquellos con el mayor número de especies son Sonora, Baja California y Chihuahua. Este trabajo representa el primer intento de integración de la información sobre la distribución geográfica de los mutílidos de México por estado.

Palabras clave: distribución geográfica; diversidad; especies; lista anotada; México

Introduction

Velvet ants belong to the family Mutillidae (Hymenoptera). Hanson & Gauld (2006) estimated that the family consists of at least 10,000 species worldwide. More recent studies suggest around 4,300 and 4,603 species around the world (Brothers & Lelej, 2017; Pagliano et al., 2020). There are eight subfamilies within the Mutillidae (Brothers & Lelej, 2017; Pagliano et al., 2020), but only Myrmosinae, Sphaeropthalminae and Mutillinae occur in the New World (Manley & Pitts, 2002). Mutilla europaea Linnaeus, 1758 was the first velvet ant described and this genus was used to encompass all the known velvet wasp species at that time. Thus, the genus Mutilla was considered the most extensive and cosmopolitan genus within Aculeata. In fact, some of the first descriptive works that included Mexican species were placed in Mutilla (Blake, 1871; 1886) or Sphaeropthalma (Blake, 1871). However, advances in the taxonomy of velvet ants have allowed many species previously recognized within Mutilla or Sphaeropthalma to be placed in other genera such as Ephuta Say, 1836; Dasymutilla Ashmead, 1899; Pseudomethoca Ashmead, 1896; and Timulla Ashmead, 1899, which occur in Mexico.

Much of the recent knowledge about the diversity of velvet ant species in Mexico comes from various types of taxonomic information, such as descriptions of new taxa or taxonomic revisions (Manley & Pitts, 2007; Pitts, 2007). Pagliano et al. (2020) attempt to integrate the diversity of the world’s velvet ants by mentioning the distribution of species at the country level. After the work of Pagliano et al. (2020), there is no integrative approach that includes an updated checklist of the velvet ants of Mexico.

Mittermeier et al. (1997), suggest that Mexico holds about 10 percent of the biodiversity of the world, at least in some groups such as mammals, birds, reptiles and plants. According to Pagliano et al. (2020), the number of velvet ants known for Mexico is 265 (plus 23 incertae sedis species and a nomen dubium). In this sense, whatever the reasons, the diversity of velvet ants occurring in Mexico seems to be underestimated. The aim of this work is to provide an integrative analysis of the velvet ants of Mexico and create the first checklist of velvet ants of the country with annotations on their distribution by state.

Materials and methods

To document the diversity of velvet ants of Mexico, the previous work on the catalogue of the velvet ants of the Neotropical region, including Mexico (Nonveiller, 1990) and that of the World (Pagliano et al., 2020) were used as basis. The checklist was generated in alphabetical order but following the classification and nomenclature of the velvet ants used by Pagliano et al. (2020). Synonyms are included. The section named “Distribution” contains the name of the state where the species occur. The new records for Mexico were marked in bold in the checklist and the incertae sedis species and a nomen dubium are shown at the end of the checklist. (See Appendix A). Two maps of Mexico were created to show the richness of genera and species for each state in a color gradient.

Two sources of information were used to generate the checklist. In the first place, specialized literature coming from general catalogues or monographs (Blake, 1871; Blake, 1886; Krombein, 1979; Nonveiller 1990) as well as different taxonomic works focused on specific groups of velvet ants, like those on Dasymutilla (Manley & Pitts, 2002; Manley & Pitts, 2007; Williams, 2023) Odontophotopsis (Pitts, 2007), Invreiella (Quintero & Cambra, 2011; Waldren et al., 2020), Sphaeropthalma (Schuster, 1958; Pitts, 2006), Acrophotopsis (Pitts & McGugh, 2002), Lomachaeta (Pitts & Manley, 2004; Williams & Pitts, 2007; Williams & Pitts, 2009; Williams et al., 2019), Lophostigma (Quintero & Cambra, 1996), Acanthophotopsis (Tanner et al., 2009), Silvorientilla (Williams et al., 2018) and Dilophotopsis (Wilson & Pitts, 2008).

The second source of information came from digitalized databases. The databases consulted were the following:

  • ASUHIC Arizona State University Hasbrouck Insect Collection

  • NMNH National Museum of Natural History, Smithsonian Institution, Washington, D.C.

  • BMNH Natural History Museum, London, UK

  • FMNH Field Museum of Natural History, Chicago, Illinois, USA

  • AMNH American Museum of Natural History, NY, USA

  • MCZ Museum of Comparative Zoology, Harvard University, Cambridge, MA

  • CAS California Academy of Sciences, San Francisco, CA

  • TTU Museum of Texas Tech University, Lubbock, TX

  • Faculty of Natural Sciences, UAQ, Querétaro, Mexico and CE-UAEM (mantained at Universidad Autónoma del Estado de Morelos, Mexico)

The database available on the web page Ecdysis Portal Home (https://ecdysis.org/) was also consulted. This web page contained the information of the following institutions:

  • PSUC Frost Entomological Museum, Pennsylvania State University, USA

  • NMSU New Mexico State Collection of Arthropods, New Mexico State University, USA

  • CSUC Gillette Museum of Arthropod Diversity, Colorado State University, USA

  • EFIC University of North Texas Elm Fork Insect Collection, University of North Texas, USA

  • DMNS Denver Museum of Nature and Science, Denver, Colorado, USA

  • WRME J. B. Wallis / R. E. Roughley Museum of Entomology University of Manitoba, Canada

  • MSUC The Albert J. Cook Arthropod Research Collection Michigan State University, USA

  • CUAC Clemson University Arthropod Collection Clemson University, USA

  • SDNHM San Diego Natural History Museum Entomology, San Diego California, USA

  • MNHUT Entomology Collection at the Natural History Museum of Utah, Salt Lake City, USA

  • NAAC Northern Arizona University - Arthropod Collection, Northern Arizona University, USA

  • UAIC University of Arizona Insect Collection, University of Arizona, USA

  • DPGNHC Dugway Proving Ground Natural History Collection, Utah, USA

  • UCSBIZC University of California Santa Barbara Invertebrate Zoology Collection, University of California Santa Barbara, USA

  • ASUHIC Arizona State University Hasbrouck Insect Collection, Arizona State University, USA

Results

A total of 369 species of velvet ants distributed in three subfamilies and 19 genera were registered. The checklist adds 106 new records compared to those obtained in Pagliano et al., (2020), which represents an increase of 40.30% of species richness. The subfamily Sphaeropthalminae was represented with 16 genera and 303 species, the Mutillinae was represented with two genera and 65 species and the Myrmosinae was represented with one genus and one species (Figs. 1-2).

Figura 1 Pie diagram showing the percentage of genera per subfamily. 

Figura 2 Pie diagram showing the percentage of species per subfamily. Note: The subfamily Myrmosinae represents 0.2%. 

The number of genera per state was highly biased, with some states much better represented than others. An extreme example is that between Sonora and Tlaxcala with 13 and zero genera respectively as shown in in the bar graph (Fig. 3) and the map that shows the number of genera per state in a color gradient (Fig. 4).

Figura 3 Bar graph showing the number of genera per state. 

Figura 4 Map of Mexico showing the number of genera per state in a color gradient. 

The number of species per state was also highly biased. For example, there were 94 species recorded for Sonora while zero species have been recorded for Tlaxcala (Figs. 5-6).

Figura 5 Bar graph showing the number of species per state. 

Figura 6 Map of Mexico showing the number of species per state in a color gradient. 

The most species rich genus was Dasymutilla with 114 species, while Myrmosula, Hoplognathoca and Pertyella were represented by one species each (Fig. 7).

Figura 7 Bar graph showing the number of species per genus. 

A total of 41 species are known for Mexico, but with no additional information on the state they occur.

Discusion

Due to its geographical location and the meeting of the Nearctic and Neotropics, Mexico is one of the so-called megadiverse countries, with many different ecosystems and a great diversity of species, making it one of the most biodiverse countries in taxonomic groups such as mammals, reptiles, amphibians and plants (CONABIO, 2008). However, with regard to the data on velvet ants obtained in this paper, this does not appear to be the case. In a study of velvet ants, Menard and Mitchell (2013) recorded 98 species in 10 genera for the state of Oklahoma, USA. This state has a very similar size to the Mexican state of Sonora with 181,037 km² and 179,355 km² respectively. These authors mention that there is a bias in the collection, as some districts are much better represented than others. They conclude that a greater effort would significantly increase the diversity of ant species in the state. Sonora was the state best represented in terms of number of genera (13) and species (94), which is very similar to Oklahoma. However, many of the species found in Sonora came from the same locality or from localities very close to each other. Some localities that occur frequently in the material studied are, for example, Álamos, Desemboque and Bahía de San Carlos, so, as in Oklahoma, there are still large unexplored regions where there are no collection efforts.

The number of species per genus correlates with other biological groups, i. e. there are some species that are numerically more dominant than others. For example, within the family Cleridae (Coleoptera), the genus Cymatodera Gray, 1832 is represented by 118 species in Mexico, while other genera such as Isoforma Opitz, 2015 are represented by one species (Araujo-Castillo et al., 2024). However, the geographical distribution of the genera and species of velvet ants found in this work differs considerably from other groups. The ants are better known in the states of Veracruz and Chiapas (Dáttilo et al., 2019), while the beetles are better known in the states of Chiapas, Oaxaca and Veracruz (Araujo-Castillo et al., 2024), in contrast to the velvet ants, which are better represented in the northern states of Mexico such as Sonora, Baja California, and Chihuahua.

Possible explanations for these patterns can be through the so-called “shortfalls” (or lack of knowledge on biodiversity) (Hortal et al., 2015). The most direct are the Linnean and Wallacean shortfalls. For example, lack of knowledge of the number of velvet ant species for Mexico (Linnean) is a consequence of those yet to be collected in remote areas, and those already collected, but in need to be described or treated taxonomically. Pagliano et al., (2020) recognized 265 species of velvet ants for Mexico, however, two of them are now synonyms, reducing the number to 263 species.

The Wallacean shortfall refers to the lack of knowledge on the geographical distribution of species. In general, this shortfall is a consequence of the influence of the capabilities and efforts of scientists, resulting in geographic biases on the distribution of species. In this regard, most of the new discoveries reported in this paper are species whose natural geographic range overlaps between some states of the United States and Mexico. Historically, there have been taxonomists in the United States who have contributed to the knowledge of this group, while Mexico was hardly known, with most of the knowledge of Mexican velvet ants coming from the studies of American taxonomists.

Another shortfall is the Hutchinsonian, which refers to the gaps of knowledge of species´life histories, functional roles and responses to habitat changes. According to the patterns of velvet ants for Mexico, it is hypothesized that the most diverse genera (Dasymutilla and Sphaeropthalma), both members of the subfamily Sphaeropthalminae, have recently diversified in the deserts of North America due to cyclical climatic changes in the Pleistocene, producing species that now live in the Chihuahuan or Sonoran Desert (Pitts et al., 2010).

A fulfilling of these gaps will enable a better understanding of the richness of the velvet ants in Mexico. Certainly, an increase in collecting activity focused on local and global inventories and the training of specialized taxonomists will lead to a much better understanding of the taxonomy and other possible studies of the Mexican velvet ants.

Acknowledgments.

I thank the academic and the executive editors of the journal, as well as the anonymous reviewers whose comments and corrections have improved this paper

Literature cited

André, E. (1898) Étude sur les Mutillides du Museum de Paris. Annales de la Société Entomologique de France, 67, 1-79. [ Links ]

André, E. (1906) Species descriptions of Mutillidae nouvelles d´Amérique. Revue d´Entomologie, 25, 32-39 [ Links ]

Araujo-Castillo, S. A., Romero-Nápoles, J. R., Rifkind, J., Flores-Dávila, M., García-Martínez, O., Cerna-Chávez, E. (2024) Biodiversidad de la familia Cleridae (Coleoptera: Cleridae) de México. Folia Entomológica Mexicana (nueva serie), 10, 20241001. https://doi.org/10.53749/fem.2024.10.01 [ Links ]

Ashmead, W. H. (1896) Descriptions of new parasitic Hymenoptera. Transactions of the American Entomological Society, 23 (2), 179-234. [ Links ]

Ashmead, W. H. (1897) Description of a new mutillid from Florida. Entomological News, 8, 20-21. [ Links ]

Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45-60. [ Links ]

Baker, C. F. (1905) New Western Mutillidae, I. Invertebrata Pacfica, 1, 93-99. [ Links ]

Bischoff, H. (1920) Monographie der Mutilliden Afrikas. Archiv für Naturgeschichte Nicolaische, Berlin, 830 pp. [ Links ]

Blake, C. A. (1870) Descriptions of two new species of Mutilla from Mexico. Transactions of the American Entomological Society, 3, 68. https://doi.org/10.2307/25076236 [ Links ]

Blake, C. A. (1871) Synopsis of the Mutillidae of North America. Transactions of the American Entomological Society, 3, 217-265. [ Links ]

Blake, C. A. (1872) Synopsis of N. Am. Mutillidae. Transactions of the American Entomological Society, 4, 71-76. [ Links ]

Blake, C. A. (1879) Catalogue of the Mutillidae of North America, with descriptions of new species. Transactions of the American Entomological Society, 7, 243-254. https://doi.org/10.2307/25076376 [ Links ]

Blake, C. A. (1886) Monograph of the Mutillidae of North America. Transactions of the American Entomological Society, 13, 179-286. https://doi.org/10.5962/bhl.title.60182 [ Links ]

Bradley, J. C. (1916) A revision of Ephuta Say, a genus of Mutillidae equivalent to the species group scrupea of Fox. Transactions of the American Entomological Society, 43, 187-198. [ Links ]

Bradley, J. C. (1917) Contributions toward a monograph of the Mutillidae and their allies of America north of Mexico. IV. A review of the Myrmosidae Transactions of the American Entomological Society, 43, 247-290. [ Links ]

Brothers, D. J. (1975) Phylogeny and classification of the aculeate Hymenoptera, with special reference to Mutillidae. University of Kansas Science Bulletin, 50, 438-648. [ Links ]

Brothers, D. J. (1982) Two new species of Mutillidae associated with Halictus hesperus (Halictidae) in Panama (Hymenoptera). Sociobiology, 7 (2), 205-212. [ Links ]

Brothers, D. J., Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1-97. https://doi.org/10.3897/jhr.60.20091 [ Links ]

Caldwell, A. H. (1939) A new species of Pseudomethoca from Arizona. The Pan-Pacific Entomologist, 15 (2), 94-95. [ Links ]

Cameron, P. (1894-1900) Fam. Mutillidae. In: Godman, F. D., Salvin, O. (eds), Biologia Centrali-Americana. Insecta. Hymenoptera. (Fossores.) Vol. II (151 pp. + 2 pls: pp. 259-328, pl. XII [1894]; pp. 329-376, pl. XIV [1895]; pp. 377-395 [1896]; p. 404 [1899]; pp. iii, v, x-xi, 405-413 [1900]). R. H. Porter, London. [ Links ]

Cameron, P. (1905) On some Hymenoptera collected by Prof. C. F. Baker in Nicaragua. Invertebrata Pacifica, 1, 99-104. [ Links ]

Casal, O. H. (1967) Comentarios sobre Acrophotopsis Schuster, con la descripción de una nueva especie de México (Hymenoptera: Mutillidae). Physis, 74, 1-4. [ Links ]

Cockerell, T. D. A. (1894) Descriptions of new Hymenoptera. Entomological News, 5, 199-201. [ Links ]

Cockerell, T. D. A. (1895) Descriptions of new Hymenoptera. Entomological News, 6, 60-64. [ Links ]

Cockerell, T. D. A. (1897) New insects from Embudo, New Mexico. Annals and Magazine of Natural History, 20, 510-515. https://doi.org/10.1080/00222939709487392 [ Links ]

Cockerell, T. D. A. (1915) New Mutillidae from California. The Entomologist, 48 (630), 249-250. [ Links ]

Cockerell, T. D. A., Casad, J. E. (1894) New species of Mutillidae. Entomological News, 5, 293-296. [ Links ]

CONABIO. (2008) Capital natural de México. vol. I: Conocimiento actual de la biodiversidad. Comisión Nacional para el Conocimiento y Uso de la Biodiversidad, México. https://doi.org/10.5962/bhl.title.118843 [ Links ]

Cresson, E. T. (1865) Descriptions of some new species of Mutilla from California. Proceedings of the Entomological Society of Philadelphia, 4, 385-390. [ Links ]

Cresson, E. T. (1875) Descriptions of new species of Mutilla. Transactions of the American Entomological Society, 5, 119-120. https://doi.org/10.2307/25076298 [ Links ]

Dalla Torre, C. G. (1897) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonimicus. Volumen VIII: Fossores (Sphegidae), 749 pp. [ Links ]

Dáttilo, W., Vásquez-Bolaños, M., Ahuatzin, D. A., Antoniazzi, R., Chávez-González, E., Corro, E., Luna, P., Guevara, R., Villalobos, F., Madrigal-Chavero, R., de Faria Falcão, J. C., Bonilla-Ramírez, A., García-Romero, A. R., De la Mora, A., Ramírez-Hernández, A., Escalante-Jiménez, A. L., Martínez-Falcón, A. P., Villarreal, A. I., García-Colón-Sandoval, A., Aponte, B., Juárez-Juárez, B., Castillo-Guevara, C., Moreno, C. E., Albor, C., Martínez-Tlapa, D. L., Huber-Sannwald, E., Escobar, F., Montiel-Reyes, F. J., Varela-Hernández, F., Castaño-Meneses, G., Pérez Lachaud, G., Pérez-Toledo, G. R., Alcalá-Martínez, I., Rivera-Salinas, I. S., Chairez-Hernández, I., Chamorro-Florescano, I. A., Hernández-Flores, J., Martínez-Toledo, J., Lachaud, J. P., Reyes-Muñoz, J. L., Valenzuela-González, J. E., Horta-Vega, J. V., Cruz-Labana, J. D., Reynoso-Campos, J. J., Navarrete-Heredia, J. L., Rodríguez-Garza, J. A., Pérez-Domínguez, J. F., Benítez-Malvido, J., Ennis, K. K., Sáenz, L., Díaz-Montiel, L. A., Tarango-Arámbula, L. A., Quiroz-Robledo, L. N., Rosas-Mejía, M., Villalvazo-Palacios, M., Gómez-Lazaga, M., Cuautle, M., Aguilar-Méndez, M. J., Baena, M. L., Madora-Astudillo, M., Rocha-Ortega, M., Pale, M., García-Martínez, M. A., Soto-Cárdenas, M. A., Correa-Ramírez, M. M., Janda, M., Rojas, P., Torres-Ricario, R., Jones, R. W., Coates, R., Gómez-Acevedo, S. L., Ugalde-Lezama, S., Philpott, S. M., Joaqui, T., Marques, T., Zamora-Gutiérrez, V., Martínez-Mandujano, V., Hajian-Forooshani, Z., MacGregor-Fors, I. (2019) Ants of Mexico: incidence and abundance along the Nearctic-Neotropical interface. Ecology, 101, e02944. 10.1002/ecy [ Links ]

Fabricius, J. C. (1775) Systema Entomologiae sistens Insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. Korte, Flensburgi et Lipsiae. 28 + 832 pp. [ Links ]

Fabricius, J. C. (1798) Supplementum Entomologiae Systematicae. Proft et Storch (Hafnia [= Copenhagen]), [I-IV,] 572 pp. [ Links ]

Ferguson, W. E. (1967) Male sphaeropthalmine mutillid wasps of the Nevada Test Site. Brigham Young University Science Bulletin, Biology Series, 8, 1-26. https://doi.org/10.5962/bhl.part.7443 [ Links ]

Fox, W. J. (1892) Description of four new species of fossorial Hymenoptera from California. Entomological News, 3, 170-172. [ Links ]

Fox, W. J. (1893) Report on some mexican Hymenoptera, principally from lower California. Proceedings of the California Academy of Sciences ser, 2 (4), 1-25. [ Links ]

Fox, W. J. (1894) Second report on some Hymenoptera from Lower California. Proceedings of the California Academy of Sciences, 4, 93-95. [ Links ]

Fox, W. J. (1899) The North American Mutillidae. Transactions of the American Entomological Society, 25, 219-292. [ Links ]

Gerstaecker, A. (1874) Mutillarum Americae meridionalis indigenarum synopsis systematica et synonymica. Archiv für Naturgeschichte, 40, 299-328. [ Links ]

Gray, G. (1832) The Class Insecta arranged by the Baron Cuvier with supplementary additions to each order by Edward Griffith and Edward Pidgeon, and notices of new genera and species by George Gray, 15, 372-373, 516. [ Links ]

Hanson, P. E., Gauld, I. D. (2006) Hymenoptera de la región neotropical. Memoirs of the American Entomological Institute. Gainesville, FL. USA. 994 pp. [ Links ]

Hortal, J., De Bello, F., Diniz-Filho, J. A. F., Lewinsohn, T. M., Lobo, J. M., Ladle, R. J. (2015) Seven shortfalls that beset large-scale knowledge of biodiversity. Annual Review of Ecology, Evolution, and Systematics, 46 (1), 523-549. https://doi.org/10.1146/annurev-ecolsys-112414-054400 [ Links ]

Howard, L. O. (1901) The Insect Book: a popular account of the bees, wasps, ants, grasshoppers, flies and other North American insects exclusive of the butterflies, moths, and beetles, with full life histories, tables and bibliographies. William Briggs, Toronto. 425 pp. https://doi.org/10.5962/bhl.title.66168 [ Links ]

Krombein, K. V. (1979) Mutillidae. In: Krombein, K. V., Hurd, P. D. Jr., Smith, D. R., Burks, B. D. (eds.). Pp 1276-1314. Catalog of Hymenoptera in America North of Mexico. Vol. 2, Apocrita (Aculeata). Smithsonian Institution Press, Washington, D.C. https://doi.org/10.1086/412023 [ Links ]

Latreille, P. A. (1802) Histoire naturelle générale et particulière des crustacés et des insectes. Tome 3. Paris: F. Dufart, xii + 467 pp. https://doi.org/10.5962/bhl.title.15764 [ Links ]

Lepeletier de Saint Fargeau, A. (1845) Histoire naturelle des insectes. Hymenoptères. Tome troisième. Roret (Paris), 646 pp. [ Links ]

Linnaeus C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines genera, species cum characteribus, differentris, synonymis, locis. Editos decima, reformata. Laurentii Salvii, Stockholm. [ Links ]

Luz, D. R., Williams, K. A., Bartholomay, P. R. (2016) The mutillid wasps of the Dasymutilla paradoxa species-group (Hymenoptera: Mutillidae). Zootaxa, 4193 (2), 361-372. https://doi.org/10.11646/zootaxa.4193.2.10 [ Links ]

Manley, D. G. (1999) Dasymutilla tomberlini, a new species of velvet ant (Hymenoptera: Mutillidae) from New Mexico. Proceedings of the Entomological Society of Washington, 101, 412-413. [ Links ]

Manley, D. G. (2003) Dasymutilla jalisco, a new species of velvet ant (Hymenoptera: Mutillidae) plus new synonymy for Dasymutilla canina (Smith). Proceedings of the Entomological Society of Washington, 105, 679-684. [ Links ]

Manley, D. G., Deyrup, M. A. (1987) A new species of Dasymutilla (Hymenoptera: Mutillidae) from Florida. Journal of Entomological Science, 22, 123-125. https://doi.org/10.18474/0749-8004-22.2.123 [ Links ]

Manley, D. G., Pitts, J. P. (2002) A key to genera and subgenera of Mutillidae (Hymenoptera) in America North of Mexico with description of a new genus. Journal of Hymenoptera Research, 11 (1), 72-100. [ Links ]

Manley, D. G. , Pitts, J. P. (2007) Tropical and subtropical velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae) with descriptions of 45 new species. Zootaxa, 1487, 1-128. https://doi.org/10.11646/zootaxa.1487.1.1 [ Links ]

Manley, D. G. , Williams, K. A., Pitts, J. P. (2020) Keys to Nearctic velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae), with notes on taxonomic changes since Krombein (1979). Proceedings of the Entomological Society of Washington, 122 (2), 335-414. https://doi.org/10.4289/0013-8797.122.2.335 [ Links ]

Melander, A. L. (1903) Notes on North American Mutillidae, with descriptions of new species. Transactions of the American Entomological Society, 29, 291-230. [ Links ]

Menard, K. L., Mitchell, J. M. (2013) Velvet ants, past and present: A county-wide checklist of the distribution and diversity of Mutillidae (Insecta: Hymenoptera) in Oklahoma including two new state records, and new behavioral observations for Dasymutilla foxi (Cockerell). Entomologica Americana, 119 (3-4), 57-79. https://doi.org/10.1664/13-ra-009.1 [ Links ]

Mickel, C. E. (1924) A revision of the mutillid wasps of the genera Myrmilloides and Pseudomethoca occurring in America North of Mexico. Proceedings of the United States National Museum, 64 (15), 1-52. https://doi.org/10.5479/si.00963801.64-2505.1 [ Links ]

Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. United States National Museum Bulletin, 143, 1-351. https://doi.org/10.5479/si.03629236.143.1 [ Links ]

Mickel, C. E. (1935) Descriptions and records of Nearctic mutillid wasps of the genera Myrmilloides and Pseudomethoca (Hymenoptera: Mutillidae). Transactions of the American Entomological Society, 61, 383-398. [ Links ]

Mickel, C. E. (1936) New species and records of Nearctic mutillid wasps of the genus Dasymutilla (Hymenoptera). Annals of the Entomological Society of America, 29, 29-60. https://doi.org/10.1093/aesa/29.1.29 [ Links ]

Mickel, C. E (1937) The mutillid wasps of the genus Timulla which occur in North America North of Mexico. Entomologica Americana, 37 (1), 1-56. [ Links ]

Mickel, C. E. (1938) The Neotropical mutillid wasps of the genus Timulla Ashmead (Hymenoptera: Mutillidae). Transactions of the Royal Entomological Society of London, 87 (22), 529-679. https://doi.org/10.1111/j.1365-2311.1938.tb00728.x [ Links ]

Mickel, C. E. (1940) Two new species of Lomachaeta, with key to described species (Hymenoptera, Mutillidae). The Pan-Pacific Entomologist, 16 (3), 127-131. [ Links ]

Mickel, C. E. (1952) The Mutillidae (Wasps) of British Guiana. Zoologica (New York), 37 (3), 105-150. https://doi.org/10.5962/p.184691 [ Links ]

Mickel, C. E. (1966) A new Dasymutilla from Baja California. The Pan-Pacific Entomologist, 42, 338-339. [ Links ]

Mickel, C. E. (1969) Pseudomethoca willei n. sp. reared from cells of the bee Lasioglossum (Dialictus) umbripenne (Ellis) (Hymenoptera: Mutillidae: Apoidea). Journal of the Kansas Entomological Society, 42 (4), 524-526. [ Links ]

Mickel, C. E., Clausen, P. E. (1983) New species of the genus Odontophotopsis subgenus Odontophotopsis Viereck (Hymenoptera; Mutillidae) of North America. Annals of the Entomological Society of America, 76, 539-551. https://doi.org/10.1093/aesa/76.3.539 [ Links ]

Mittermeier, R. A., Goettsch-Mittermeier, C., P. Robles-Gil., Wilson, E. O. (1997) Megadiversidad: los países biológicamente más ricos del mundo. Cemex-Agrupación Sierra Madre, México. [ Links ]

Nonveiller, G. (1990) Catalogue of the Mutillidae, Myrmosidae and Bradynobaenidae of the Neotropical region including Mexico (Insecta: Hymenoptera). Hymenopterorum Catalogus (nova editio), Pars 18 (150 pp.). SPB Academic Publishing, The Hague [ Links ]

Opitz, W. (2015) Systematics of the checkered beetle subfamily Platynopterinae Spinola (Coleoptera, Cleridae). Linzer Biologische Beiträge, 47 (2), 1513-1572. [ Links ]

Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P. L., Williams, K. A., Romano, M. (2020) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bolletino del Museo Regionale di Scienze Naturali, 36 (1-2), 5-425. [ Links ]

Pitts, J, P. (2005) Description of a new species in the Sphaeropthalma pensylvanica species-group (Hymenoptera: Mutillidae). Zootaxa, 1050 (1), 39-43. https://doi.org/10.11646/zootaxa.1050.4 [ Links ]

Pitts, J, P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1-20. https://doi.org/10.11646/zootaxa.1248.1.1 [ Links ]

Pitts, J. P. (2007) Revision of the Odontophotopsis Viereck (Hymenoptera; Mutillidae) Part I, with a description of a new genus Laminatilla. Zootaxa, 1619 (1), 1-43. https://doi.org/10.11646/zootaxa.1619.1.1 [ Links ]

Pitts, J. P ., Manley, D. G. (2002) Description of females of Stethophotopsis Pitts and Sphaeropthalma (Photopsioides Schuster) (Hymenoptera: Mutillidae). Proceedings of the Entomological Society of Washington, 104 (3), 672-679. [ Links ]

Pitts, J. P., Manley, D. G. (2004) A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474, 1-27. https://doi.org/10.11646/zootaxa.474.1.1 [ Links ]

Pitts, J, P., McHugh, J, V. (2000) Stethophotopsis, a new genus of Sphaeropthalmini (Mutillidae: Sphaeropthalminae) with a brachypterous male from Arizona. Journal of Hymenoptera Research, 9 (1), 29-33. [ Links ]

Pitts, J, P., McHugh, J, V. (2002) Revision of Acrophotopsis (Mutillidae: Sphaeropthalminae), with a new species from Baja California. Journal of Hymenoptera Research, 11 (2), 338-349. [ Links ]

Pitts, J. P. , Wilson, J. S., von Dohlen, C. D. (2010) Evolution of the nocturnal Nearctic Sphaeropthalminae velvet ants (Hymenoptera: Mutillidae) driven by Neogene orogeny and Pleistocene glaciation. Molecular Phylogenetics and Evolution, 56, 134-145. [ Links ]

Provancher, L. (1882) Faune entomologique du Canada traitent des Hyménoptères. Le Naturaliste Canadien 12, 4-51. [ Links ]

Provancher, L. (1887) Additions et corrections à la faune hyménoptérologique de la province de Quebec. [Quebec?: s. n] 475 pp. https://doi.org/10.5962/bhl.title.38326 [ Links ]

Quintero, A. D., Cambra, R. A. (1996) The Mexican and Central American species of Lophostigma Mickel, including a new species, new distribution records, and taxonomic notes for the genus (Hymenoptera: Mutillidae). The Pan-Pacific Entomologist, 72 (2), 92-101. [ Links ]

Quintero, A. D., Cambra, R. (2011) Revision of the Mexican endemic genus Invreiella Suárez (Hymenoptera: Mutillidae), with the description of a new species. Transactions of the American Entomological Society, 13 (1+2), 217-224. https://doi.org/10.3157/061.137.0114 [ Links ]

Radoszkowski, O. (1861) Descriptions of several new species of Hymenoptera. Horae Societatis Entomologicae Rossicae, 1, 79-86. [ Links ]

Rohwer, S. A. (1909) New Hymenoptera from Western United States. Transactions of the American Entomological Society, 35, 99-136. [ Links ]

Rohwer, S. A. (1910) Some new wasps from New Jersey. Proceedings of the Entomological Society of Washington, 12, 49-52. [ Links ]

Rohwer, S. A. (1912) Descriptions of new species of wasps in the collections of the United States National Museum. Proceedings of the United States National Museum, 41, 447-478. https://doi.org/10.5479/si.00963801.41-1869.447 [ Links ]

Saussure, H. de (1867) Mutillarum novarum species aliquot. Annales de la Societé Entomologique de France, 7, 351-364. [ Links ]

Say, T. (1836) Descriptions of new species of North American Hymenoptera, and observation on some already described. Boston Journal of Natural History, 1 (3), 209-305. [ Links ]

Schuster, R. M. (1945) An interesting new brachypterous species of Photopsis (Hymenoptera: Mutillidae). The Pan-Pacific Entomlogist, 21 (4), 149-151. [ Links ]

Schuster, R. M. (1949) Contributions toward a monograph of the Mutillidae of the Neotropical region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, 29 (3-4), 59-140. [ Links ]

Schuster, R. M. (1951) A revision of the genus Ephuta (Mutillidae) in America North of Mexico. New York Entomological Society, 59 (1), 1-43. [ Links ]

Schuster, R. M. (1952) Notes on some North American Mutillidae I. Some new species of Odontophotopsis. Bulletin of the Brooklyn Entomological Society, 47, 43-49. [ Links ]

Schuster, R. M. (1958) A revision of the Sphaeropthalminae, Mutillidae of America north of Mexico. II. Entomologica Americana (n. s.), 37, 1-30. [ Links ]

Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Taylor and Francis, London, 206 pp. [ Links ]

Smith F. (1862) Descriptions of new species of aculeate Hymenoptera, collected at Panama by R. W. Stretch, Esq, with a list of described species, and the various localities where they have previously occurred. Transactions of the Entomological Society of London, 3 (1), 29-44. [ Links ]

Smith F. (1879) Descriptions of New Species of Hymenoptera in the Collection of the British Museum. Printed by order of the Trustees, London. https://doi.org/10.5962/bhl.title.60089 [ Links ]

Suárez, F. J. (1962) Datos sobre Mutílidos neotropicales. III. Un nuevo género de Centroamérica y países noroccidentales de América del Sur (Hymenoptera). Eos, Revista Española de Entomología Madrid, 38 (1), 115-146. [ Links ]

Suárez, F. J. (1966 [“1965”]) Datos sobre Mutílidos neotropicales. VI. Dos nuevos géneros de Pseudomethocini de Méjico y Brasil (Hymenoptera). Eos, Revista Española de Entomología Madrid, 41 (2-3), 471-482. [ Links ]

Tanner, D. A., Boehme, N. F., Pitts, J. P. (2009) Review of Acanthophotopsis Schuster (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 18 (2), 192-204. [ Links ]

Viereck, H. L. (1903) Hymenoptera from southern California and New Mexico, with descriptions of new species. Proceedings of the Academy of Natural Sciences, Philadelphia, 54, 737-743. [ Links ]

Viereck, H. L. (1904) The species of Odontophotopsis (Hymenoptera). Transactions of the American Entomological Society, 30, 81-92. [ Links ]

Viereck, H. L. (1906) Notes and descriptions of Hymenoptera from the western United States. Transactions of the American Entomological Society, 32 (2), 173-247. [ Links ]

Viereck, H. L. (1924) Descriptions of new Hymenoptera in the Canadian National Collection, Ottawa. The Canadian Entomologist, 56 (5), 110-112. https://doi.org/10.4039/ent56110-5 [ Links ]

Waldren, G. C., Williams, K. A. , Cambra, R. A., Pitts, J. P . (2020) Systematic revision of the North American velvet ant genus Invreiella Suárez (Hymenoptera: Mutillidae) with description of eleven new species. Zootaxa, 4894 (2), 151-205. https://doi.org/10.11646/zootaxa.4894.2.1 [ Links ]

Wasbauer, M.S. (1973) Some new taxa in the Myrmosinae with keys to the females in North America (Hymenoptera: Tiphiidae). The Pan-Pacific Entomologist, 49 (4), 325-337. [ Links ]

Williams, K. A. (2023) Taxonomic updates for diurnal velvet ants (Hymenoptera: Mutillidae) in the United States of America. Zootaxa, 5301 (1), 105-123. https://doi.org/10.11646/zootaxa.5301.1.5 [ Links ]

Williams, K. A. , Cambra, R. A , Bartholomay, P. R., Luz, D. R., Quintero, D., Pitts, J. P. (2019) Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa, 4564 (1), 101-136. https://doi.org/10.11646/zootaxa.4564.1.4 [ Links ]

Williams, K. A. , Cambra, R. A., Waldren, G. C. , Quintero, D. , Pitts, J. P. (2019[“2018”]) Silvorientilla, gen. nov. A new neotropical velvet ant genus (Hymenoptera: Mutillidae). Folia Entomológica Mexicana (nueva serie), 4 (3), 91-109. [ Links ]

Williams, K. A. , Pitts, J. P . (2007) New species of the predominately temperate velvet ant genera Lomachaeta Mickel and Sphaeropthalma Blake from Central and northern South America (Hymenoptera: Mutillidae) Transactions of the American Entomological Society, 133 (3), 297-326. https://doi.org/10.3157/0002-8320-133.3.297 [ Links ]

Williams, K. A. , Pitts, J. P. (2008) Three species masquerading as one: updating the taxonomy of Pseudomethoca russeola Mickel and P. donaeanae (Cockerell & Fox) (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 17 (2), 127-133. [ Links ]

Williams, K. A. , Pitts, J. P. (2009) Eight new species of Lomachaeta Mickel and the synonymy of Smicromutilla Mickel (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 18 (2), 227-243. [ Links ]

Wilson, J. S., Pitts, J. M. (2008) Revision of velvet ant genus Dilophotopsis Schuster (Hymenoptera: Mutillidae) by using molecular and morphological data, with implications for desert biogeography. Annals of the Entomological Society of America, 101 (3), 514-524. https://doi.org/10.1603/0013-8746(2008)101[514:rovagd]2.0.co;2 [ Links ]

How to cite: Varela-Hernández, F. (2025) The velvet ants (Hymenoptera, Mutillidae) of Mexico: a taxonomic account. Acta Zoológica Mexicana (nueva serie), 41, 1-55.

Responsible editor: Magdalena Cruz Rosales

Appendix A

Checklist of the velvet ants of Mexico

Family Mutillidae Latreille, 1802

Subfamily Mutillinae Latreille, 1802

Tribe Mutillini Latreille, 1802

Genus Ephuta Say, 1836

1. Ephuta catriona (Cameron, 1894)

Mutilla catriona Cameron, 1894.

Mutilla blakei Cameron, 1894.

Distribution: Guerrero.

2. Ephuta cinaloa (Blake, 1871) .

Mutilla cinaloa Blake, 1871.

Mutilla sinaloa Cameron, 1894.

Distribution: Morelos, Nayarit.

3. Ephuta copano (Blake, 1871) .

Mutilla copano Blake, 1871.

Mutilla susura Melander, 1903.

Distribution: Mexico.

4. Ephuta culta (Cameron, 1894) .

Mutilla culta Cameron, 1894.

Distribution: Tabasco.

5. Ephuta dioetus (Cameron, 1896) .

Sphaerophthalma dioetus Cameron, 1896.

Distribution: Mexico.

6. Ephuta ecarinata Schuster, 1951.

Distribution: Mexico.

7. Ephuta herberti (Cameron, 1894).

Mutilla herberti, Cameron, 1894.

Distribution: Mexico.

8. Ephuta idiastia (Cameron, 1894).

Mutilla idiastia Cameron, 1894.

Distribution: Guerrero, Sonora.

9. Ephuta megaera (Cameron, 1894).

Mutilla megaera Cameron, 1894.

Distribution: Guerrero.

10. Ephuta pomona (Cameron, 1894).

Mutilla pomona Cameron, 1894.

Distribution: Guerrero, Yucatán.

11. Ephuta sapiens (Smith, 1879).

Mutilla sapiens Smith, 1879.

Mutilla astuta Cameron, 1894.

Distribution: Veracruz.

12. Ephuta teapae (Cameron, 1894) .

Mutilla teapae Cameron, 1894.

Distribution: Tabasco.

13. Ephuta tegulica Bradley, 1916.

Distribution: Mexico.

14. Ephuta tolteca (Blake, 1871).

Mutilla tolteca Blake, 1871.

Distribution: Mexico.

15. Ephuta yucatana (Blake, 1871).

Mutilla yucatana Blake, 1871.

Distribution: Campeche.

Tribu Trogaspidiini Bischoff, 1920

Genus Timulla Ashmead, 1899

16. Timulla adrastis Mickel, 1938.

Timulla pilatrix Mickel, 1938.

Distribution: Jalisco, Michoacán, Morelos, Nayarit.

17. Timulla aerolla (Cameron, 1894) .

Mutilla aerolla Cameron, 1894.

Distribution: Guerrero.

18. Timulla asperata Mickel, 1938.

Distribution: Guerrero.

19. Timulla collina Mickel, 1938.

Distribution: Coahuila, Colima.

20. Timulla cyllene (Cameron 1894) .

Mutilla cyllene Cameron, 1894.

Distribution: Guerrero, Jalisco, Morelos, Nayarit, San Luis Potosí, Veracruz.

21. Timulla cyra (Cameron, 1894) .

Mutilla cyra Cameron, 1894.

Mutilla vanduara Cameron, 1894.

Distribution: Guerrero, Morelos.

22. Timulla daedala (Cameron, 1894) .

Mutilla daedala Cameron, 1894.

Distribution: Yucatán.

23. Timulla damalis Mickel, 1938.

Distribution: Veracruz. .

24. Timulla delecta Mickel, 1938.

Distribution: Tabasco, Veracruz.

25. Timulla derelicta (Smith, 1879) .

Mutilla derelicta Smith, 1879.

Distribution: Mexico.

26. Timulla direpta Mickel, 1938.

Distribution: Durango, Tabasco, Veracruz.

27. Timulla disgrega Mickel, 1938.

Distribution: Veracruz.

28. Timulla divergens Mickel, 1938.

Distribution: Coahuila, Oaxaca.

29. Timulla diversita Mickel, 1938.

Distribution: Chiapas.

30. Timulla dubitata Mickel, 1938.

Mutilla dubitata Smith, 1855.

Mutilla dubiatta [sic.] Blake, 1886.

Timulla (Timulla) dubitata fugitiva Mickel, 1937.

Timulla (Timulla) murcia Mickel, 1938.

Distribution: Sonora.

31.Timulla falsa (Dalla Torre, 1897).

Mutilla falsa Dalla Torre, 1897.

Mutilla rufofemorata Cameron, 1894.

Distribution: Guerrero, Morelos.

32. Timulla ferocula Mickel, 1938.

Distribution: Yucatan.

33. Timulla gaumeri (Cameron, 1894).

Mutilla gaumeri Cameron, 1894.

Distribution: Veracruz, Yucatán.

34. Timulla grotei (Blake, 1871).

Mutilla grotei Blake, 1871.

Mutilla fulviventris Gerstaecker, 1874.

Distribution: Chihuahua, Durango, Jalisco, Zacatecas.

35. Timulla hoegei (Cameron, 1894).

Mutilla hoegei Cameron, 1894.

Distribution: Veracruz.

36. Timulla impudica (Cameron, 1894).

Mutilla impudica Cameron, 1894.

Distribution: Guerrero.

37. Timulla incensita Mickel, 1938.

Distribution: Oaxaca, Veracruz.

38. Timulla interstincta Mickel, 1938.

Distribution: Mexico.

39. Timulla leona (Blake, 1871).

Mutilla leona Blake, 1871.

Distribution: Guerrero, Nuevo León, San Luis Potosí, Zacatecas.

40. Timulla lilea (Cameron 1894).

Mutilla lilea Cameron, 1894.

Mutilla orthona (Cameron, 1894).

Distribution: Jalisco.

41. Timulla mexicana (Cameron, 1894).

Mutilla mexicana Cameron, 1894.

Mutila amulae Cameron, 1894.

Distribution: Guerrero, Morelos, Oaxaca.

42. Timulla navasota (Bradley, 1916) .

Mutilla navasota Bradley, 1916.

Timulla navasota nebulosa Mickel, 1937.

Distribution: Chihuahua, Coahuila, Guanajuato, Nuevo León, Tamaulipas.

43. Timulla neobule Mickel, 1937.

Timulla nicholi Mickel, 1937.

Distribution: Jalisco, Morelos, Sinaloa.

44. Timulla nitela Mickel, 1937.

Distribution: Nayarit.

45. Timulla oajaca (Blake, 1871).

Mutilla oajaca Blake, 1871.

Mutilla ardens Gerstaecker, 1874.

Mutilla mazatlanae Cameron, 1894.

Mutilla nestor Fox, 1899.

Mutilla ornata Howard, 1901.

Distribution: Chihuahua, Estado de México, Jalisco, Morelos, Nayarit, Sinaloa.

46. Timulla obscurella Mickel, 1938.

Distribution: Guerrero, Veracruz.

47. Timulla operosa Mickel, 1938.

Distribution: Jalisco, Sonora.

48. Timulla ordinaria (Smith, 1879) .

Mutilla ordinaria Smith, 1879.

Distribution: Chiapas.

49. Timulla osberti (Cameron, 1894) .

Mutilla osberti Cameron, 1894.

Distribution: Tabasco, Veracruz.

50. Timulla philyra Mickel, 1938.

Distribution: Mexico.

51. Timulla porcata (Cameron, 1894) .

Mutilla porcata Cameron, 1894.

Timulla phiala Mickel, 1938.

Timulla bituberculata Mickel, 1938.

Distribution: Oaxaca.

52. Timulla prema (Cameron, 1894) .

Mutilla prema Cameron 1894.

Distribution: Coahuila, Guerrero.

53. Timulla prominens (Cameron, 1894) .

Mutilla prominens Cameron, 1894.

Distribution: Guerrero, Jalisco, Nayarit, Morelos, Oaxaca, San Luis Potosí, Sinaloa, Sonora.

54. Timulla psyra (Cameron, 1894) .

Mutilla psyra Cameron, 1894.

Distribution: Sinaloa, San Luis Potosí, Yucatán.

55. Timulla rhanis Mickel, 1938.

Distribution: Aguascalientes, Jalisco, Morelos.

56. Timulla ruida Mickel, 1938.

Distribution: Nayarit.

57. Timulla saundersi Mickel, 1938.

Distribution: Veracruz.

58. Timulla simplex Mickel, 1938.

Distribution: Chiapas, Jalisco.

59. Timulla subrobusta (Cameron, 1894) .

Mutilla talus Cameron, 1894.

Mutilla subrobusta Cameron, 1894.

Mutilla clutha Cameron, 1894.

Distribution: San Luis Potosí.

60. Timulla suffusa Mickel, 1938.

Distribution: Mexico.

61. Timulla suspensa (Gerstaecker, 1874).

Mutilla suspensa Gerstaecker, 1874.

Timulla suspensa jonesi Mickel, 1937.

Timulla suspensa sonora Mickel, 1937.

Distribution: Baja California, Chihuahua, Durango, Nuevo León.

62. Timulla taygete Mickel, 1938.

Timulla aureata Mickel, 1938.

Distribution: Oaxaca, Veracruz, Yucatán.

63. Timulla trifolia Mickel, 1938.

Distribution: Yucatán.

64. Timulla tyro Mickel, 1937.

Distribution: Nayarit.

65. Timulla vagans (Fabricius, 1798) .

Mutilla vagans Fabricius, 1798.

Mutilla hexagona Say, 1836.

Mutilla ornativentris Cresson, 1865.

Mutilla briaxus Blake, 1871.

Mutilla canadensis Provancher, 1887.

Mutilla secunda Dalla Torre, 1897.

Distribution: Mexico.

Subfamilia Sphaeropthalminae Schuster, 1949 (1903)

Tribu Dasymutillini Brothers & Lelej, 2017

Genus Dasymutilla Ashmead, 1899

66. Dasymutilla albiceris Mickel, 1936.

Distribution: Baja California.

67. Dasymutilla altamira (Blake, 1871)

Mutilla altamira Blake, 1871.

Sphaerophthalma [sic.] laothoe Cameron, 1895.

Distribution: Chiapas, Guerrero, Oaxaca, Jalisco.

68. Dasymutilla Andréniformis (Smith, 1855) .

Mutilla Andréniformis Smith, 1855.

Distribution: Veracruz.

69. Dasymutilla apicalata (Blake, 1871) .

Mutilla apicalata Blake, 1871.

Sphaerophthalma guerreroensis Cameron, 1895.

Sphaerophthalma [sic.] perfidiosa Cameron, 1895.

Distribution: Baja California Sur, Chiapas, Chihuahua, Durango, Guerrero, Jalisco, Michoacán, Nayarit, Oaxaca, San Luis Potosí, Sinaloa, Sonora, Veracruz.

70. Dasymutilla arachnoides (Smith, 1855) .

Mutilla arachnoides Smith, 1855.

Sphaerophthalma [sic.] hiera Cameron, 1895.

Sphaerophthalma [sic.] ephippiata Cameron, 1895.

Distribution: Campeche, Chiapas, Chihuahua, Colima, Guerrero; Jalisco, Morelos, Nayarit, Oaxaca, Puebla, Querétaro, Quintana Roo, San Luis Potosí, Tamaulipas, Veracruz, Yucatán.

71. Dasymutilla araneoides (Smith, 1862) .

Sphaerophthalma [sic.] temaxensis Cameron, 1895.

Sphaerophthalma [sic.] anthracia Cameron, 1895.

Sphaerophthalma [sic.] melanogaster Cameron, 1895.

Sphaerophthalma [sic.] waterhousei Cameron, 1895.

Dasymutilla deyrollesi Mickel, 1937.

Distribution: Campeche, Guerrero, Jalisco, Nayarit, Oaxaca, Sinaloa, Tamaulipas, Quintana Roo, Veracruz, Yucatán.

72. Dasymutilla arenivaga Mickel, 1928.

Dasymutilla megalophthalma Mickel, 1928.

Distribution: Baja California, Hidalgo, Sonora.

73. Dasymutilla argentea Manley & Pitts 2007.

Distribution: Campeche.

74. Dasymutilla asteria Mickel, 1936.

Distribution: Jalisco, Nayarit, Sonora, Sinaloa.

75. Dasymutilla atrata Manley & Pitts, 2007.

Distribution: Campeche.

76. Dasymutilla atricauda Mickel, 1936.

Distribution: Baja California, Sonora.

77. Dasymutilla aureola (Cresson, 1865) .

Mutilla aureola Cresson, 1865.

Distribution: Baja California.

78. Dasymutilla azteca (Blake, 1871) .

Mutilla azteca Blake, 1871.

Distribution: Campeche.

79. Dasymutilla baleygi Manley & Pitts, 2007.

Distribution: Jalisco.

80. Dasymutilla batui Manley & Pitts, 2007.

Distribution: Hidalgo, Nayarit, Oaxaca.

81. Dasymutilla bioculata (Cresson, 1865) .

Mutilla fenestrata Lepeletier, 1845.

Mutilla bioculata Cresson, 1865.

Mutilla aegina Cresson, 1865.

Mutilla bellona Cresson, 1865.

Mutilla creusa Cresson, 1865.

Mutilla medea Cresson, 1865.

Mutilla chiron Blake, 1872.

Mutilla ursula texana Cresson, 1875.

Mutilla lepeletieri Fox, 1899.

Mutilla pyrrhus Fox, 1899.

Mutilla chlamydata Melander, 1903.

Dasymutilla vierecki Rohwer, 1910.

Dasymutilla ferrugata balabetei Rohwer, 1912.

Dasymutilla georgiana Rohwer, 1912.

Dasymutilla plesia Rohwer, 1912.

Dasymutilla balbatei Rohwer, 1912.

Dasymutilla melanippe Mickel, 1928.

Dasymutilla lauta Mickel, 1928.

Dasymutilla praegrandis Mickel, 1928.

Dasymutilla sulcatulla Mickel, 1928.

Dasymutilla russala Mickel, 1928.

Dasymutilla conformis Mickel, 1928.

Dasymutilla bimaculata Manley & Pitts, 2007.

Distribution: Baja California, Chihuahua, Coahuila, Durango, Puebla, San Luis Potosí, Sonora.

82. Dasymutilla boharti, Manley & Pitts, 2007.

Distribution: Sonora.

83. Dasymutilla bonita Mickel, 1928.

Dasymutilla poliothrix Mickel, 1928.

Distribution: Guerrero, Jalisco, Nayarit, Sinaloa, Sonora.

84. Dasymutilla buenavista Manley & Pitts, 2007.

Distribution: Campeche.

85. Dasymutilla californica (Radoszkowski, 1861) .

Mutilla californica Radoszkowski, 1861.

Dasymutilla abdita Mickel, 1928.

Distribution: Baja California, Baja California Sur.

86. Dasymutilla campogrande Manley & Pitts, 2007.

Distribution: Mexico.

87. Dasymutilla canina (Smith, 1855) .

Mutilla canina Smith, 1855.

Mutilla sumichrasti Saussure, 1867.

Sphaerophthalma [sic.] philina Cameron, 1895.

Mutilla transmarina Dalla Torre, 1897.

Mutilla variicolor André, 1898.

Distribution: Chiapas, Guerrero, Jalisco, Morelos, Sinaloa, Oaxaca, Puebla.

88. Dasymutilla cardinalis Mickel, 1937.

Distribution: Oaxaca, Veracruz.

89. Dasymutilla chilcotti Manley & Pitts, 2007.

Distribution: San Luis Potosí.

90. Dasymutilla chionothrix Manley & Pitts, 2007.

Dasymutilla truxali Manley & Pitts, 2007.

Distribution: Morelos, Sonora.

91. Dasymutilla cirrhomeris Manley & Pitts, 2007.

Distribution: Sinaloa.

92. Dasymutilla clytemnestra (Fox, 1899) .

Mutilla clytemenstra Fox, 1899.

Distribution: Baja California.

93. Dasymutilla coccineohirta (Blake, 1871) .

Mutilla coccineohirta Blake, 1871.

Mutilla ochracea Blake, 1879.

Sphaerophthalma [sic.] venifica Blake, 1886.

Mutilla progne Fox, 1899.

Dasymutilla aletina Cockerell, 1915.

Distribution: Baja California.

94. Dasymutilla combusta (Smith, 1855).

Mutilla combusta Smith, 1855.

Distribution: Hidalgo, Jalisco, San Luis Potosí, Veracruz.

95. Dasymutilla connectens (Cameron, 1895) .

Sphaerophthalma [sic.] connectens Cameron, 1895.

Dasymutilla helva Mickel, 1928.

Dasymutilla ocydrome Mickel, 1928.

Distribution: Baja California, Baja California Sur, Chihuahua, Jalisco, Sinaloa, Sonora.

96. Dasymutilla cressoni (Blake, 1870).

Mutilla cressoni Blake, 1870.

Distribution: Chiapas, Chihuahua, Colima Guerrero, Jalisco, Morelos, Michoacán, Nayarit, Oaxaca, Puebla, Sinaloa, Veracruz.

97. Dasymutilla dammersi Mickel, 1936.

Distribution: Baja California Sur.

98. Dasymutilla eminentia Mickel, 1928.

Distribution: Baja California Sur, San Luis Potosí, Sinaloa, Sonora.

99. Dasymutilla erythrina (Say, 1836) .

Mutilla erythrina Say, 1836.

Dasymutilla fulgida Mickel, 1928.

Dasymutilla pallida Mickel, 1928.

Distribution: Aguascalientes, Chiapas, Chihuahua, Ciudad de Mexico. Colima, Durango, Guanajuato, Guerrero, Jalisco, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, Sinaloa, Sonora, Veracruz, Yucatán.

100. Dasymutilla eurynome Mickel, 1928.

Distribution: Nayarit.

101. Dasymutilla fasciventris Mickel, 1938.

Dasymutilla chamela Manley & Pitts, 2007.

Dasymutilla citromaculosa Manley & Pitts, 2007.

Distribution: Jalisco, Morelos, Michoacán, Nayarit, Sinaloa, Puebla, Sonora.

102. Dasymutilla fasciventroides Manley et al. 2020.

Distribution: Jalisco, Nayarit, Sinaloa, Sonora.

103. Dasymutilla ferruginea (Smith, 1879).

Mutilla ferruginea (Smith, 1879).

Dasymutilla chrysocoma Mickel, 1928.

Distribution: Chihuahua, Colima, Guerrero, Jalisco, Michoacán, Nayarit, Oaxaca, Sinaloa.

104. Dasymutilla fimbriata Manley & Pitts, 2007.

Distribution: Guerrero, Michoacán.

105. Dasymutilla flammifera Mickel, 1928.

Distribution: Baja California.

106. Dasymutilla formicalia Rohwer, 1912.

Distribution: Ciudad de Mexico, Hidalgo, Jalisco, Morelos, Sinaloa.

107. Dasymutilla formosa (Blake, 1871).

Mutilla formosa Blake, 1871.

Mutilla batopilas Blake, 1871.

Mutilla batophila Dalla Torre, 1897.

Sphaeropthalma batopilas Cameron, 1895.

Distribution: Aguascalientes, Chiapas, Guanajuato, Jalisco, Michoacán, Nayarit, Oaxaca, Querétaro, Sinaloa, Sonora, Veracruz, Yucatán.

108. Dasymutilla foxi (Cockerell, 1894) .

Mutilla foxi Cockerell, 1894.

Mutilla phoenix Fox, 1899.

Spaherophthalma [sic.] heterochroa Cockerell & Casad, 1894.

Spaherophthalma [sic.] dugesii Cockerell & Casad, 1894.

Spaherophthalma [sic.] foxi Cockerell, 1894.

Distribution: Aguascalientes, Baja California, Baja California Sur, Coahuila, Chihuahua, Guerrero, Guanajuato, Jalisco, Michoacán, Nayarit, Nuevo León, San Luis Potosí, Sonora, Sinaloa, Tamaulipas, Zacatecas.

109. Dasymutilla gloriosa (Saussure, 1867) .

Mutilla gloriosa (Saussure, 1867) .

Dasymutilla reperticia Mickel, 1928.

Dasymutilla chisos Mickel, 1928.

Distribution: Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Guanajuato, Nayarit, Nuevo León, San Luis Potosí, Sinaloa, Sonora.

110. Dasymutilla gorgon (Blake, 1871) .

Mutilla gorgon Blake, 1871.

Distribution: Mexico.

111. Dasymutilla griswoldi Manley & Pitts, 2007.

Distribution: Chihuahua, Jalisco.

112. Dasymutilla holotricha (Saussure, 1867) .

Mutilla holotricha Saussure, 1867.

Distribution: Morelos, Nayarit, Oaxaca.

113. Dasymutilla imperialis Manley & Pitts, 2007.

Distribution: Baja California Sur.

114. Dasymutilla impressa Manley & Pitts, 2007.

Distribution: Jalisco, Michoacán, Nayarit, Sinaloa, Sonora.

115. Dasymutilla iztapa (Blake, 1871) .

Mutilla iztapa (Blake, 1871) .

Sphaerophthalma [sic.] rodriguezi Cameron, 1895.

Distribution: Campeche, Chiapas, Guerrero, Jalisco, Oaxaca, San Luis Potosí, Sinaloa, Veracruz.

116. Dasymutilla jalisco Manley, 2003.

Distribution: Chiapas, Guerrero, Jalisco, Morelos.

117. Dasymutilla klugii (Gray, 1832) .

Mutilla klugii Gray, 1832.

Mutilla orcus Cresson, 1865.

Sphaerophthalma [sic.] klugi Cameron, 1895.

Distribution: Ciudad de Mexico, Chihuahua, Coahuila, Durango, Guanajuato, Hidalgo, Jalisco, Nuevo León, Puebla, Querétaro, Sonora, San Luis Potosí, Tamaulipas.

118. Dasymutilla lachesis Mickel, 1966.

Distribution: Baja California, Baja California Sur, Veracruz.

119. Dasymutilla latebalteata (Cameron, 1897) .

Sphaerophthalma [sic.] latebalteata Cameron, 1897.

Dasymutilla veracruz Manley & Pitts, 2007.

Distribution: Veracruz.

120. Dasymutilla leve Manley & Pitts, 2007.

Distribution: Sinaloa.

121. Dasymutilla loreto Manley & Pitts, 2007.

Distribution: Baja California Sur.

122. Dasymutilla magna (Cresson, 1865) .

Mutilla magna (Cresson, 1865) .

Distribution: Baja California, Baja California Sur, Sonora, Sinaloa, San Luis Potosí, Tamaulipas.

123. Dasymutilla magnifica Mickel, 1928.

Distribution: Baja California, Campeche, Chiapas, Chihuahua, San Luis Potosí, Sinaloa, Sonora, Veracruz.

124. Dasymutilla mazatlanorum Manley & Pitts, 2007.

Distribution: Sinaloa, Sonora.

125. Dasymutilla melanargyrea (André, 1898) .

Mutilla melanargyrea André, 1898.

Distribution: Morelos, Oaxaca, Puebla.

126. Dasymutilla modgudi Manley & Pitts, 2007.

Distribution: Guerrero.

127. Dasymutilla monticola (Cresson, 1865) .

Mutilla caneo Blake, 1879.

Mutilla mixtura Blake, 1879.

Mutilla myrrha Fox, 1899.

Mutilla monticola Cresson, 1865.

Mutilla eximia Blake, 1886.

Ephuta boulderensis Rohwer, 1909.

Distribution: Tamaulipas.

128. Dasymutilla monstrosa Manley & Pitts, 2007.

Distribution: Chihuahua, Chiapas, Colima, Jalisco, Nayarit, Sinaloa, Sonora, Querétaro, Zacatecas.

129. Dasymutilla munifica (Smith, 1879) .

Mutilla munifica Smith, 1879.

Distribution: Chiapas, San Luis Potosí, Veracruz, Yucatán.

130. Dasymutilla myrice Mickel, 1928.

Distribution: Chihuahua.

131. Dasymutilla naranjo Manley & Pitts, 2007.

Distribution: Veracruz.

132. Dasymutilla niphopilis Manley & Pitts, 2007.

Distribution: Baja California Sur, Chiapas, Hidalgo, Querétaro, Sonora.

133. Dasymutilla nocticaro Manley & Pitts, 2007.

Dasymutilla concordia Manley & Pitts, 2007.

Distribution: Chihuahua, Nayarit, Sinaloa, Sonora, Zacatecas.

134. Dasymutilla nocturna Mickel, 1928.

Dasymutilla subhyalina Mickel, 1928.

Dasymutilla paranocturna Barr & Hurd, 1947.

Distribution: Sonora.

135. Dasymutilla nogalensis Mickel, 1928.

Distribution: Chihuahua, Coahuila, Sinaloa, Sonora.

136. Dasymutilla ocydrome Mickel, 1928.

Distribution: Baja California Sur, Nayarit.

137. Dasymutilla ocyrae (Cameron, 1895) .

Sphaerophthalma [sic.] ocyrae Cameron, 1895.

Dasymutilla gaumeri Mickel, 1937.

Distribution: Campeche, Chiapas, Yucatán.

138. Dasymutilla pallene (Cameron, 1895) .

Sphaerophthalma [sic.] pallene Cameron, 1895.

Distribution: Durango, Sinaloa, Sonora.

139. Dasymutilla paradoxa (Gerstaecker, 1874) .

Mutilla paradoxa Gerstaecker, 1874.

Dasymutilla guanacaste Manley & Pitts, 2007.

Sphaeropthalma icaris (Cameron, 1895) .

Distribution: Veracruz.

140. Dasymutilla paraparadoxa Luz, Bartholomay & Williams, 2016.

Distribution: Quintana Roo.

141. Dasymutilla parkerorum Manley & Pitts, 2007.

Distribution: Campeche, Quintana Roo, Yucatán.

142. Dasymutilla personata (Cameron, 1895) .

Spaherophthalma [sic.] personata Cameron, 1895.

Mutilla sumichrasti Saussure, 1867.

Distribution: Jalisco, Morelos, Nayarit, Oaxaca, Sinaloa.

143. Dasymutilla perilla Mickel, 1928.

Distribution: Coahuila, Nuevo León, Tamaulipas.

144. Dasymutilla phaon (Fox, 1899) .

Mutilla phaon Fox, 1899.

Distribution: Baja California, Chihuahua, Sinaloa.

145. Dasymutilla phya (Cameron, 1895) .

Sphaerophthalma [sic.] phya Cameron, 1895.

Sphaerophthalma [sic.] melissa Cameron, 1895.

Sphaeropthalma [sic.] sleipniri Manley & Pitts, 2007.

Distribution: Quintana Roo.

146. Dasymutilla proclea (Cameron, 1895) .

Sphaerophthalma [sic.] proclea Cameron, 1895.

Sphaerophthalma [sic.] cobira Cameron, 1895.

Distribution: Campeche, Yucatán.

147. Dasymutilla pseudopappus (Cockerell, 1895) .

Sphaerophthalma pseudopappus Cockerell, 1895.

Distribution: Baja California, Baja California Sur, Sinaloa, Sonora, Zacatecas.

148. Dasymutilla pulchra (Smith, 1855) .

Mutilla pulchra Smith, 1855.

Mutilla nortoni Blake, 1870.

Mutilla ducalis Smith, 1879.

Sphaerophthalma [sic.] chiron Cameron, 1897.

Distribution: Baja California, Campeche, Chiapas, Colima, Chihuahua, Estado de México, Guerrero, Jalisco, Morelos, Nayarit, Oaxaca, Querétaro, Quintana Roo, Sonora, Sinaloa, Tabasco, Veracruz, Yucatán.

149. Dasymutilla relata (Cameron, 1895)

Sphaerophthalma [sic.] realta Cameron, 1895

Distribution: Yucatán

150. Dasymutilla rufipilis Manley & Pitts, 2007

Distribution: Puebla

151. Dasymutilla rutiliventris (André, 1898)

Mutilla rutiliventris André, 1898

Distribution: Jalisco, Sonora

152. Dasymutilla sackenii (Cresson, 1865)

Mutilla sackenii Cresson, 1865

Mutilla erudita Cresson, 1875

Distribution: Baja California, Baja California Sur, Sonora

153. Dasymutilla saetigera Mickel, 1928

Dasymutilla polia Mickel, 1928

Distribution: Sinaloa, Sonora

154. Dasymutilla satanas Mickel, 1928

Dasymutilla mimula Mickel, 1928

Distribution: Baja California, Baja California Sur, Nuevo León, Sonora

155. Dasymutilla scaber (Fox, 1894)

Sphaerophthalma [sic.] scaber Fox, 1894

Distribution: Baja California Sur

156. Dasymutilla schumanni (Cameron, 1895)

Sphaerophthalma [sic.] schumanni Cameron, 1895

Distribution: Nayarit

157. Dasymutilla serenitas Mickel, 1928

Distribution: Coahuila

158. Dasymutilla sicheliana (Saussure, 1867)

Mutilla sicheliana Saussure, 1868

Sphaerophthalma [sic.] prunotincta Cockerell, 1895

Sphaerophthalma [sic.] thera Cameron, 1895

Sphaerophthalma [sic.] thalia Cameron, 1895

Mutilla gynaecologica Dalla Torre, 1897

Dasymutilla intermixta Mickel, 1928

Distribution: Aguascalientes, Chihuahua, Colima, Durango, Guerrero, Jalisco, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Sinaloa, Sonora, Veracruz, Zacatecas

159. Dasymutilla sinaloa Manley & Pitts, 2007

Distribution: Baja California Sur, Sinaloa

160. Dasymutilla snowrum (Cockerell, 1897)

Sphaerophthalma [sic.] snowrum Cockerell, 1897

Mutilla poecilonota Melander, 1903

Distribution: Chihuahua, Coahuila

161. Dasymutilla sophrona Mickel, 1928

Distribution: Morelos, San Luis Potosí, Sinaloa, Sonora

162. Dasymutilla spilota Manley & Pitts, 2007

Dasymutilla ionothorax Manley & Pitts, 2007

Distribution: Colima, Nayarit

163. Dasymutilla stevensi Mickel, 1928

Dasymutilla medora Mickel, 1928

Distribution: Chihuahua, Coahuila, Guanajuato, Sinaloa

164. Dasymutilla terminata (Smith, 1879)

Mutilla terminata Smith, 1879

Mutilla sumichrasti Saussure, 1867

Distribution: Oaxaca, Zacatecas

165. Dasymutilla toluca (Blake, 1871)

Mutilla (Sphaeropthalma) toluca Blake, 1871

Mutilla (Sphaeropthalma) ariadne Blake, 1871

Mutilla (Sphaeropthalma) solola Blake, 1871

Sphaerophthalma [sic.] postica Cameron, 1897

Distribution: Chiapas, Colima, Nayarit, Jalisco, Veracruz, Guerrero, Campeche, Oaxaca

166. Dasymutilla tomasi Manley & Pitts, 2007

Distribution: Baja California Sur

167. Dasymutilla tomberlini Manley, 1999

Distribution: Sonora

168. Dasymutilla twegeni Manley & Pitts, 2007

Distribution: Quintana Roo, Yucatán

169. Dasymutilla ulkei (Cresson, 1865)

Mutilla ulkei Cresson, 1865

Distribution: Baja California Sur

170. Dasymutilla vesta (Cresson, 1865)

Mutilla vesta Cresson, 1865

Scolia unicincta Provancher, 1882

Mutilla monozona Dalla Torre, 1897

Mutilla sappho Fox, 1899

Mutilla agenor Fox, 1899

Mutilla zella Rohwer, 1910

Pycnomutilla harmoniiformis Rohwer, 1912

Dasymutilla errans Rohwer, 1912

Dasymutilla bosquensis Rohwer, 1912

Dasymutilla ferrugatella Rohwer, 1912

Dasymutilla coloradella Rohwer, 1912

Dasymutilla coloradella virginica Rohwer, 1912

Dasymutilla coloradella kamloopsensis Rohwer, 1912

Dasymutilla texensis Rohwer, 1912

Dasymutilla mesillae Rohwer, 1912

Dasymutilla carolina Rohwer, 1912

Dasymutilla columbiana Mickel, 1928

Distribution: Chihuahua, Tamaulipas

171. Dasymutilla vestita (Lepeletier, 1845)

Mutilla vestita Lepeletier, 1845

Mutilla montezumae Lepeletier, 1845

Mutilla fulvohirta Cresson, 1865

Sphaerophthalma [sic.] townsendi Cockerell, 1894

Sphaerophthalma [sic.] aspasia Cameron, 1895

Mutilla aspasioides Dalla Torre, 1897

Ephuta californica var. euchroa Cockerell, 1897

Dasymutilla vandala Mickel, 1928

Dasymutilla cotulla Mickel, 1928

Distribution: Aguascalientes, Chihuahua, Ciudad de Mexico, Coahuila, Colima, Durango, Hidalgo, Jalisco, Nuevo León, Oaxaca, Querétaro, San Luis Potosí, Sonora, Tamaulipas, Zacatecas

172. Dasymutilla vulpina (Smith, 1879)

Mutilla vulpina Smith, 1879 Oaxaca

Distribution: Ciudad de Mexico, Colima, Durango, Estado de Mexico, Guerrero, Jalisco, Morelos, Nayarit, Sinaloa, Oaxaca, Puebla

173. Dasymutilla waco (Blake, 1871)

Mutilla (Sphaeropthalma) waco Blake, 1871

Distribution: Coahuila, Nuevo León, San Luis Potosí

174. Dasymutilla wasbaueri Manley & Pitts, 2007

Distribution: Guerrero, Jalisco, Nayarit, Nuevo León, Oaxaca, San Luis Potosí, Tamaulipas, Veracruz

175. Dasymutilla whartoni Manley & Pitts, 2007

Distribution: Guerrero

176. Dasymutilla xalisco (Blake, 1871)

Mutilla (Sphaeropthalma) xalisco Blake, 1871

Distribution: Colima, Jalisco, Nayarit, Oaxaca

177. Dasymutilla yucatana (Cameron, 1895)

Sphaerophthalma [sic.] yucatana Cameron, 1895

Mutilla yucatanicola Dalla Torre, 1897

Distribution: Yucatán

178. Dasymutilla zelaya (Blake, 1871)

Mutilla (Sphaeropthalma) zelaya Blake, 1871

Distribution: Chihuahua, Coahuila, Durango

179. Dasymutilla zoster Manley & Pitts, 2007

Distribution: Chiapas, Oaxaca

Genus Lomachaeta Mickel, 1936

180. Lomachaeta chionothrix Pitts & Manley, 2004

Distribution: Jalisco, Nayarit

181. Lomachaeta cirrhomeris Pitts & Manley, 2004

Distribution: Baja California, Sonora

182. Lomachaeta crocopinna Pitts & Manley, 2004

Distribution: Sonora

183. Lomachaeta eironeia Williams, 2019

Distribution: Sonora

184. Lomachaeta hedera Williams & Pitts, 2009

Distribution: Baja California Sur

185. Lomachaeta hicksi Mickel, 1936

Lomachaeta hicksi Mickel, 1936

Lomachaeta coloradensis Mickel, 1936

Lomachaeta punctinota Mickel, 1936

Lomachaeta minutula Mickel, 1936

Lomachaeta variegata Mickel, 1940

Lomachaeta formosula Mickel, 1940

Dasymutilla mickeli Manley & Deyrup, 1987

Distribution: Coahuila

186. Lomachaeta litosisyra Williams & Pitts, 2009

Distribution: Sonora

187. Lomachaeta polemomechana Williams & Pitts, 2009

Distribution: Sonora

188. Lomachaeta ptilohyalus Pitts & Manley, 2004

Distribution: Oaxaca

189. Lomachaeta theresa Williams & Pitts, 2009

Distribution: Sonora

190. Lomachaeta vacamuerta Williams & Pitts, 2009

Distribution: Sonora

Tribu Pseudomethocini Brothers, 1975

Genus Hoplognathoca Suárez, 1962

191. Hoplognathoca robinsoni (Blake, 1871)

Mutilla robinsonii Blake, 1871

Mutilla croesus Blake, 1871

Sphaerophthalma [sic.] respublicana Cameron, 1894

Distribution: Veracruz

Genus Invreiella Suárez, 1966

192. Invreiella acuminata Waldren, 2020

Distribution: Mexico

193. Invreiella australis Waldren, 2020

Distribution: Chiapas, Oaxaca

194. Invreiella bimaculata Waldren, 2020

Distribution: Sinaloa

195. Invreiella breviclypeata Waldren, 2020

Distribution: Sinaloa

196. Invreiella cardinalis (Gerstaecker, 1874)

Mutilla cardinalis Gerstaecker, 1874

Sphaerophthalma [sic.] jocularis Cameron, 1894

Sphaerophthalma [sic.] megacantha Cockerell & Casad, 1894

Invreiella curoei Quintero & Cambra, 2011

Distribution: Veracruz, Durango, Estado de Mexico. Guanajuato, Guerrero, Jalisco, Morelos, Oaxaca, Puebla, Zacatecas

197. Invreiella cephalargia (Mickel, 1924)

Pseudomethoca cephalargia Mickel, 1924

Distribution: Chihuahua

198. Invreiella cuernavaca Waldren, 2020

Distribution: Morelos

199. Invreiella chihuahuensis Waldren, 2020

Distribution: Chihuahua, Durango

200. Invreiella erythrocephala Waldren, 2020

Distribution: Morelos

201. Invreiella manleyi Waldren, 2020

Distribution: Chihuahua

202. Invreiella mesomexicana Waldren, 2020

Distribution: Guanajuato, Jalisco, Nayarit, Zacatecas

203. Invreiella satrapa (Gerstaecker, 1874)

Mutilla satrapa Gerstaecker, 1874

Mutilla macrocephala Smith, 1879

Distribution: Coahuila, Durango, Estado de Mexico, Guanajuato, Hidalgo, Ciudad de Mexico, Michoacán, San Luis Potosí, Querétaro, Veracruz, Zacatecas

204. Invreiella suarezi Waldren, 2020

Distribution: Guerrero

205. Invreiella tequila Waldren, 2020

Distribution: Jalisco, Morelos

Genus Lophostigma Mickel, 1952

206. Lophostigma grisselli Cambra & Quintero, 1996

Distribution: Yucatán

207. Lophostigma subgracilis (Cameron, 1895)

Sphaerophthalma [sic.] subgracilis Cameron, 1895

Distribution: Tabasco

Genus Pertyella Mickel, 1952

208. Pertyella salutatrix (Smith, 1879)

Mutilla salutatrix Smith, 1879

Distribution: Veracruz

Genus Pseudomethoca Ashmead, 1896

209. Pseudomethoca ajattara Williams & Pitts, 2008

Distribution: Estado de Mexico

210. Pseudomethoca anthracina (Fox, 1892)

Sphaerophthalma [sic.] anthracina Fox, 1892

Mutilla anthracicolor Dalla Torre, 1897

Mutilla harpalyce Fox, 1899

Distribution: Chihuahua, Durango

211. Pseudomethoca athamas (Fox, 1899)

Mutilla athamas Fox, 1899

Distribution: Baja California Sur

212. Pseudomethoca auripes Blake, 1871

Distribution: Mexico

213. Pseudomethoca bequaerti Mickel, 1924

Distribution: Chihuahua, Sonora

214. Pseudomethoca bisignata (Blake, 1871)

Mutilla bisignata Blake, 1871

Distribution: Mexico

215. Pseudomethoca brazoria (Blake, 1871)

Mutilla brazoria, Blake, 1871

Distribution: Coahuila

216. Pseudomethoca chiapa (Blake, 1871)

Mutilla chiapa Blake, 1871

Distribution: Mexico

217. Pseudomethoca chontalensis (Cameron, 1895)

Sphaerophthalma [sic.] chontalensis Cameron, 1895

Sphaerophthalma [sic.] cephalica Cameron, 1895

Mutilla guatemalensis Dalla Torre, 1897

Distribution: Veracruz

218. Pseudomethoca connectens (Cresson, 1865)

Mutilla conncetens Cresson, 1865

Mutilla eggeri Dalla Torre, 1897

Pseudomethoca helva Mickel, 1928

Sphaerophthalma [sic.] connectens Cresson, 1865

Distribution: Baja California, Baja California Sur, San Luis Potosí

219. Pseudomethoca contumax (Cresson, 1865)

Mutilla contumax Cresson, 1865

Mutilla micrphthalma Gerstaecker, 1874

Distribution: Aguascalientes, Coahuila, Nuevo León

220. Pseudomethoca contumeliosa Mickel, 1935

Distribution: Baja California Sur

221. Pseudomethoca damia (Cameron, 1894)

Sphaerophthalma [sic.] damia Cameron, 1894

Distribution: Guerrero

222. Pseudomethoca donaeanae (Cockerell, 1897)

Sphaerophthalma donaeanae Cockerell, 1897

Distribution: Baja California

223. Pseudomethoca gila (Blake, 1871)

Mutilla gila Blake, 1871

Mutilla ilione Fox, 1899

Mutilla aprica Melander, 1903

Distribution: Mexico

224. Pseudomethoca gothica (Blake, 1871)

Mutilla gothica Blake, 1871

Distribution: Mexico

225. Pseudomethoca intrepida (Cameron, 1895)

Sphaerophthalma [sic.] intrepida Cameron, 1895

Distribution: Tabasco

226. Pseudomethoca izucar (Blake, 1871)

Mutilla izucar Blake, 1871

Distribution: Mexico

227. Pseudomethoca laticeps (Blake, 1871)

Mutilla laticeps Blake, 1871

Distribution: Mexico

228. Pseudomethoca leoporina (Gerstaecker, 1874)

Mutilla leoporina Gerstaecker, 1874

Distribution: Mexico

229. Pseudomethoca lerma (Blake, 1871)

Mutilla leoporina Gerstaecker, 1874

Distribution: Mexico

230. Pseudomethoca minutissima (Blake, 1871)

Mutilla minutissima Blake, 1871

Distribution: Mexico

231. Pseudomethoca mulaiki Mickel, 1938

Distribution: Tamaulipas

232. Pseudomethoca nephele (Fox, 1899)

Mutilla nephele Fox, 1899

Distribution: Colima

233. Pseudomethoca petricola (Blake, 1871)

Mutilla petricola Blake, 1871

Distribution: Puebla

234. Pseudomethoca phedyma (Cameron, 1894)

Sphaerophthalma [sic.] phedyma Cameron, 1894

Distribution: Chiapas

235. Pseudomethoca praeclara (Blake, 1886)

Sphaerophthalma [sic.] praeclara Blake, 1886

Mutilla aegaeon Fox, 1899

Nomiaephagus acuum Cockerell, 1915

Distribution: Baja California, Sonora

236. Pseudomethoca propinqua (Cresson, 1865)

Mutilla propinqua Cresson, 1865

Mutilla montivaga Cresson, 1865

Pseudomethoca sacatona Caldwell, 1939

Distribution: Chihuahua

237. Pseudomethoca psammodroma (Blake, 1871)

Mutilla psammadroma Blake, 1871

Distribution: Jalisco

238. Pseudomethoca ravula (Cameron, 1894)

Sphaerophthalma [sic.] ravula Cameron, 1894

Distribution: Chiapas, Guerrero, Jalisco, Michoacán, Morelos, Nayarit, Sinaloa, Oaxaca.

239. Pseudomethoca toumeyi (Fox, 1894)

Sphaerophthalma [sic.] toumeyi Fox, 1894

Distribution: Sonora

240. Pseudomethoca verticalis (Smith, 1855)

Mutilla verticalis Smith, 1855

Sphaerophthalma [sic.] cordovensis Cameron, 1895

Sphaerophthalma [sic.] mirandillensis Cameron, 1895

Sphaerophthalma [sic.] caroli Cameron, 1905

Distribution: Veracruz

241. Pseudomethoca volatilis (Cameron, 1895)

Sphaerophthalma [sic.] volatilis Cameron, 1895

Mutilla volans Dalla Torre, 1897

Distribution: Tabasco

242. Pseudomethoca vulnerifrons (André, 1906)

Ephuta vulnerifrons André, 1906

Distribution: Campeche

243. Pseudomethoca willei Mickel, 1969

Pseudomethoca transversa Brothers, 1982

Distribution: Oaxaca, Veracruz

244. Pseudomethoca xalapa Blake, 1871

Sphaeropthalma xalapa Blake, 1871

Distribution: Mexico

245. Pseudomethoca zapoteca (Blake, 1871)

Sphaeropthalma zapoteca Blake, 1871

Distribution: Mexico

Tribu Sphaeropthalmini Schuster, 1949 (1903)

Genus Acanthophotopsis Schuster, 1958

246. Acanthophotopsis bequaertii Schuster, 1958

Distribution: Chihuahua, Durango

247. Acanthophotopsis evansii Schuster, 1958

Distribution: Chihuahua, Coahuila, Durango

248. Acanthophotopsis falciformis Schuster 1958

Distribution: Sonora, Zacatecas

249. Acanthophotopsis snellingi Tanner & Pitts, 2009

Distribution: Chihuahua, Zacatecas

Genus Acrophotopsis Schuster, 1958

250. Acrophotopsis bergi Casal, 1967

Distribution: Chihuahua, Morelos, Puebla

551. Acrophotopsis campylognatha Schuster, 1958

Distribution: Baja California, Baja California Sur

252. Acrophotopsis dirce (Fox, 1899)

Mutilla dirce Fox, 1899

Acrophotopsis eurygnathus Schuster, 1958

Distribution: Nayarit, Sonora

253. Acrophotopsis mickeli Pitts, 2002

Distribution: Baja California, Baja California Sur

Genus Dilophotopsis Schuster, 1958

254. Dilophotopsis concolor (Cresson, 1865)

Mutilla concolor (Cresson, 1865)

Mutilla pygmaea Blake, 1879

Mutilla nanula Dalla Torre, 1897

Odontophotopsis alamonis Viereck, 1904

Distribution: Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Nayarit, San Luis Potosí, Sonora

255. Dilophotopsis paron (Cameron, 1896)

Sphaerophthalma [sic.] paron Cameron, 1896

Dilophotopsis concolor sonorensis Schuster, 1958

Distribution: Baja California, Baja California Sur

Genus Laminatilla Pitts, 2007

256. Laminatilla bicornigera (Schuster, 1958)

Distribution: Baja California Sur, Jalisco, Morelos

257. Laminatilla lamellifera (Schuster, 1958)

Odontophotopsis lamellifera Schuster, 1958

Distribution: Sonora

258. Laminatilla mixtoensis (Schuster, 1958)

Odontophotopsis mixtoensis Schuster, 1958

Distribution: Guerrero, Puebla

Genus Odontophotopsis Viereck, 1903

259. Odontophotopsis acmaea Viereck, 1904

Mutilla grata Melander, 1903

Odontophotopsis acmaeus Viereck, 1904

Distribution: Baja California, Baja California Sur, Chihuahua, Sonora

260. Odontophotopsis appacheorum Schuster, 1958

Distribution: Coahuila.

261. Odontophotopsis armata Schuster, 1958

Distribution: Baja California, Baja California Sur

262. Odontophotopsis biramosa Schuster, 1952

Distribution: Sonora

263. Odontophotopsis braccata Schuster, 1958

Distribution: Chihuahua

264. Odontophotopsis carinata Mickel, 1983

Distribution: Mexico

265. Odontophotopsis clypeata Schuster, 1958

Distribution: Baja California Sur, Durango, Sonora

266. Odontophotopsis conifera Schuster, 1958

Distribution: Sonora

267. Odontophotopsis cookie Baker, 1905

Distribution: Baja California, Baja California Sur

268. Odontophotopsis delodonta Viereck, 1904

Distribution: Sonora

269. Odontophotopsis denticoxa Mickel, 1983

Distribution: Sonora

270. Odontophotopsis dentifera Schuster, 1958

Distribution: Baja California, Baja California Sur, Sonora

271. Odontophotopsis erebus (Melander, 1903)

Mutilla erebus Melander, 1903

Mutilla avellanus Viereck, 1904

Distribution: Chihuahua, Coahuila, Tamaulipas

272. Odontophotopsis eubule (Cameron, 1896)

Spaherphthalma eubule Cameron, 1896

Mutilla hamata Melander, 1903

Distribution: Sonora

273. Odontophotopsis hammetti Pitts, 2010

Distribution: Baja California, Sonora

274. Odontophotopsis hexadonta Schuster, 1958

Distribution: Baja California

275. Odontophotopsis ignacio Schuster, 1958

Distribution: Baja California, Baja California Sur

276. Odontophotopsis inconspicua (Blake, 1886)

Photopsis inconspicuus Blake, 186

Mutilla infelix Dalla Torre, 1897

Distribution: Baja California, Baja California Sur, Sonora

277. Odontophotopsis mariae (Cameron, 1896)

Sphaerophthalma [sic.] mariae Cameron, 1896

Distribution: Nayarit

278. Odontophotopsis melicausa (Blake, 1871)

Agama melicausa Balke, 1871

Mutilla brevicornis Fox, 1899

Distribution: Baja California, Baja California Sur, Coahuila, Chihuahua, Nuevo León, Sonora

279. Odontophotopsis mexicana Pitts, 2007

Distribution: Guerrero, Jalisco

280. Odontophotopsis microdonta Ferguson, 1967

Distribution: Baja California Sur

281. Odontophotopsis multidentata Mickel, 1983

Distribution: Mexico

282. Odontophotopsis obliqua Viereck, 1924

Odontophotopsis obliquus Viereck, 1925

Distribution: Baja California

283. Odontophotopsis parva Schuster, 1958

Distribution: Sonora

284. Odontophotopsis pima Schuster, 1958

Distribution: Baja California

285. Odontophotopsis polis Pitts, 2007

Distribution: Baja California

286. Odontophotopsis quadridentata Schuster, 1958

Distribution: Baja California

287. Odontophotopsis quadrispinosa Schuster, 1958

Distribution: Baja California, Baja California Sur, Sonora

288. Odontophotopsis rubriventris (Schuster, 1958)

Sphaeropthalma facilis rubriventris Schuster, 1958

Distribution: Aguascalientes, Chihuahua, Durango, Tamaulipas

289. Odontophotopsis cerca Viereck, 1904

Odontophotopsis sercus Viereck, 1904

Distribution: Baja California, Baja California Sur

290. Odontophotopsis setifera Schuster, 1952

Distribution: Baja California

291. Odontophotopsis sonora (Schuster, 1958)

Sphaeropthalma sonora Schuster, 1958

Distribution: Baja California, Sonora

292. Odontophotopsis succinea Viereck, 1903

Odontophotopsis succineus Viereck, 1903

Odontophotopsis annulatus Baker, 1905

Distribution: Baja California

293. Odontophotopsis tenuiptera Schuster, 1958

Distribution: Chihuahua, Sinaloa, Sonora

294. Odontophotopsis territa (Cockerell, 1894)

Photopsis territus Cockerell, 1894

Distribution: Chihuahua

295. Odontophotopsis unicornis Schuster, 1958

Odontophotopsis unicornis Schuster, 1958

Distribution: Sonora

296. Odontophotopsis villosa Mickel, 1983

Distribution: Sonora

Genus Photomorphus Viereck, 1903

297. Photomorphus bicolor (Blake, 1879)

Agama bicolor Blake, 1879

Mutilla madejskii Dalla Torre, 1897

Distribution: Chihuahua, Sonora

298. Photomorphus clandestinus (Viereck, 1903)

Odontophotopsis clandestinus Viereck, 1903

Mutilla hebes Melander, 1903

Photomorphus taeniatus Schuster, 1958

Photomorphus parksi Schuster, 1958

Distribution: Coahuila, Nuevo León, Oaxaca, Sonora, Tamaulipas

299. Photomorphus cobabi Schuster, 1958

Distribution: Baja California, Morelos, Sonora

300. Photomorphus cochisae Schuster, 1958

Distribution: Mexico

301. Photomorphus comobabi Schuster, 1958

Distribution: Mexico

302. Photomorphus dentistoma Schuster, 1958

Distribution: Mexico

303. Photomorphus gamelia (Cameron, 1895)

Sphaerophthalma [sic.] gamelia Cameron, 1895

Distribution: Mexico

304. Photomorphus gamelioides Schuster, 1958

Distribution: Mexico

305. Photomorphus juanita Schuster, 1952

Photomorphus (Photomorphina) juanita Schuster, 1952

Distribution: Jalisco, Tamaulipas

306. Photomorphus mexicanella Schuster, 1952

Distribution: Baja California Sur

307. Photomorphus orizaba (Blake, 1871)

Agama orizaba Blake, 1871

Distribution: Mexico

308. Photomorphus piceogaster (Schuster, 1958)

Odontophotopsis piceogaster Schuster, 1958

Distribution: Mexico

309. Photomorphus quadriangulatus Schuster, 1958

Distribution: Mexico

310. Photomorphus sarpedon (Fox, 1899)

Mutilla sarpedon Fox, 1899

Distribution: Mexico

Genus Silvorientilla Williams & Cambra, 2019

311. Silvorientilla bonus Williams & Cambra, 2019

Distribution: Quintana Roo

312. Silvorientilla philobeddoe Williams & Cambra, 2019

Distribution: Veracruz

313. Silvorientilla prosarasoror Williams & Cambra, 2019

Distribution: Oaxaca

314. Silvorientilla separata (Smith, 1879)

Mutilla separata Smith, 1879

Distribution: Veracruz

Genus Sphaeropthalma Blake, 1871

315. Sphaeropthalma abbreviata Schuster, 1958

Distribution: Baja California

316. Sphaeropthalma abdominalis (Blake, 1886)

Photopsis abdominalis Blake, 1886

Mutilla coloradensis Dalla Torre, 1897

Distribution: Chihuahua

317. Sphaeropthalma alsvid Williams & Pitts, 2007

Distribution: Chiapas

318. Sphaeropthalma amphion (Fox, 1899)

Mutilla amphion Fox, 1899

Photopsis abstrusa Baker, 1905

Photopsis nudatus Baker, 1905

Distribution: Baja California, Baja California Sur, Sonora

319. Sphaeropthalma arota (Cresson, 1875)

Mutilla arota Cresson, 1875

Mutilla helicaon Fox, 1899

Photopsis lingulatus Viereck, 1903

Photopsis carinata Schuster, 1958

Photopsis diegueno Schuster, 1958

Photopsis coahuilae Schuster, 1958

Distribution: Baja California, Baja California Sur, Chihuahua, San Luis Potosí, Sonora, Zacatecas

320. Sphaeropthalma arvak Williams & Pitts, 2007

Distribution: Chiapas

321. Sphaeropthalma baboquivari Schuster, 1958

Distribution: Sonora

322. Sphaeropthalma bellerophon (Fox, 1899)

Mutilla bellerophon Fox, 1899

Distribution: Chihuahua

323. Sphaeropthalma blakeii (Fox, 1893)

Photopsis blakeii Fox, 1893

Mutilla gautschii Dalla Torre, 1897

Mutilla ceyx Fox, 1899

Distribution: Baja California, Baja California Sur, Sonora

324. Sphaeropthalma brachyptera (Schuster, 1945)

Photopsis brachyptera Schuster, 1945

Photopsis yavapai Schuster, 1958

Distribution: Baja California Sur, Chihuahua, Coahuila, Sonora

325. Sphaeropthalma ceyxoides Schuster, 1958

Distribution: Baja, California, Sonora

326. Sphaeropthalma chupacabra Williams & Pitts, 2007

Distribution: Chiapas

327. Sphaeropthalma clara (Cresson, 1865)

Mutilla clara Cresson, 1865

Distribution: Zacatecas

328. Sphaeropthalma coaequalis Cameron, 1896

Mutilla albicincta Fox, 1899

Mutilla piceoceps Schuster, 1958

Mutilla flavipes Schuster, 1958

Distribution: Sonora

329. Sphaeropthalma danaus (Blake, 1871)

Agama danaus Blake, 1871

Mutilla danaus Dalla Torre, 1897

Distribution: Mexico

330. Sphaeropthalma difficilis (Baker, 1905)

Photopsis difficilis Baker, 1905

Photopsis maricopella Schuster, 1958

Photopsis castanea Schuster, 1958

Photopsis fuscatella Schuster, 1958

Photopsis purismella Schuster, 1958

Photopsis parrasia Schuster, 1958

Photopsis quijotoa Schuster, 1958

Photopsis californiense Schuster, 1958

Distribution: Baja California, Baja California Sur, Chihuahua, Sonora

331. Sphaeropthalma edwardsii (Cresson, 1875)

Mutilla edwardsii Cresson, 1875

Sphaeropthalma flammifera Schuster, 1958

Distribution: Baja California

332. Sphaeropthalma facilis (Cameron, 1896)

Sphaerophthalma [sic.] facilis Cameron, 1896

Mutilla hubbardii Fox, 1899

Mutilla piceogaster Schuster, 1958

Distribution: Baja California Sur, Chihuahua, Durango, Zacatecas

333. Sphaeropthalma fuscipes Schuster, 1958

Distribution: Durango

334. Sphaeropthalma gullinbursti Williams & Pitts, 2007

Distribution: Quintana Roo

335. Sphaeropthalma hati Williams & Pitts, 2007

Distribution: Quintana Roo, Yucatán

336. Sphaeropthalma hialina (Blake, 1871)

Agama hialina Blake, 1871

Agama minuta Blake, 1872

Distribution: Baja California Sur, Chihuahua

337. Sphaeropthalma ignacio Schuster, 1958

Distribution: Baja California, Baja California Sur

338. Sphaeropthalma imperialis (Blake, 1871)

Agama imperialis Blake, 1871

Mutilla erato Blake, 1879

Distribution: Coahuila, Nuevo León, Tamaulipas

339. Sphaeropthalma insignis (Baker, 1905)

Photopsis insignis Baker, 1905

Distribution: Baja California, Baja California Sur

340. Sphaeropthalma isolatrix (Schuster, 1958)

Xenophotopsis isolatrix Schuster, 1958

Distribution: Baja California, Baja California Sur

341. Sphaeropthalma jacala Schuster, 1958

Distribution: Coahuila, Hidalgo, Jalisco, Puebla

342. Sphaeropthalma juxta (Blake, 1872)

Agama juxta Blake, 1872

Distribution: Chihuahua, Durango

343. Sphaeropthalma marpesia (Blake, 1879)

Mutilla marpesia Blake, 1879

Sphaerophthalma [sic.] luteola Blake, 1886

Mutilla imperialiformis Viereck, 1906

Sphaeropthalma (Photopsis) maricopae Schuster, 1958

Distribution: Chiapas, Baja California

344. Sphaeropthalma mendica (Blake, 1871)

Agama mendica Blake, 1871

Mutilla aspasia Blake, 1879

Photopsis nebulosus Blake, 1886

Distribution: Mexico

345. Sphaeropthalma militaris Schuster, 1958

Distribution: Baja California, Sonora

346. Sphaeropthalma nana (Ashmead, 1896)

Photopsis nana Ashmead, 1896

Mutilla acontia (Fox, 1899)

Mutilla ashmeadi Fox, 1899

Distribution: Baja California, Baja California Sur, Chihuahua

347. Sphaeropthalma nigriventris (Fox, 1893)

Photopsis nigriventris Fox, 1893

Distribution: Baja California Sur

348. Spaheropthalma nigrobalteata (Cameron, 1896)

Brachycistis exornatus Cameron, 1899

Distribution: Mexico

349. Spaheropthalma nocticaro Pitts, 2005

Distribution: Oaxaca

350. Sphaeropthalma noctivaga (Melander, 1903)

Mutilla noctivaga Melander, 1903

Mutilla infuscata Schuster, 1958

Distribution: Chihuahua, Sonora, Zacatecas

351. Sphaeropthalma nokomis (Blake, 1871)

Agama nokomis Blake, 1871

Distribution: Chihuahua

352. Sphaeropthalma orestes (Fox, 1899)

Mutilla orestes (Fox, 1899)

Mutilla pattersonae Melander, 1903

Photopsis uniformis Baker, 1905

Photopsis pedatus Baker, 1905

Photopsis ingenuus Baker, 1905

Mutilla fresnoensis Schuster, 1958

Mutilla oregana Schuster, 1958

Distribution: Baja California, Baja California Sur

353. Sphaeropthalma pallida (Blake, 1871)

Agama pallida Blake, 1871

Sphaeropthalma arizonae Schuster, 1958

Distribution: Chihuahua, Sonora

354. Sphaeropthalma pallidipes Schuster, 1958

Sphaeropthalma gila Schuster, 1958

Distribution: Sonora, Chihuahua

355. Sphaeropthalma parapenalis Ferguson, 1967

Distribution: Chihuahua

356. Sphaeropthalma pateli Schuster, 1958

Distribution: Sonora

357. Sphaeropthalma pinalea Schuster, 1958

Sphaeropthalma (Photopsis) pinalea Schuster, 1958

Distribution: Baja California Sur, Coahuila, Durango, Puebla

358. Sphaeropthalma pinales Schuster, 1958

Distribution: Mexico

359. Sphaeropthalma purissima Schuster, 1958

Photopsis purissima Schuster, 1958

Distribution: Baja California, Baja California Sur

360. Sphaeropthalma skinfaxi Williams &Pitts, 2007

Distribution: Chiapas, Oaxaca

361. Sphaeropthalma subtriangularis Schuster, 1958

Distribution: Baja California, Baja California Sur

362. Sphaeropthalma tetracuspis Schuster, 1958

Distribution: Baja California, Baja California Sur

363. Sphaeropthalma triangularis (Blake, 1871)

Agama triangularis Blake, 1871

Distribution: Chihuahua, Sonora

364. Sphaeropthalma unicolor (Cresson, 1865)

Mutilla unicolor Cresson, 1865

Mutilla phaedra Blake, 1879

Mutilla auraria Blake, 1879

Agama rustica Blake, 1879

Mutilla monochroa Dalla Torre, 1897

Sphaerophthalma [sic.] anthophora Ashmead, 1897

Dasymutilla sumneriella Cockerell, 1915

Sphaeropthalma ocellaria Schuster, 1958

Distribution: Baja California

365. Sphaeropthalma uro (Blake, 1879)

Agama uro Blake, 1879

Photopsis melanderi Baker, 1905

Sphaeropthalma stenognatha Schuster, 1958

Distribution: Coahuila, Chihuahua, Durango, Zacatecas

366. Sphaeropthalma yumaella Schuster, 1958

Sphaeropthalma (Micromutilla) yumaella Schuster, 1958

Distribution: Baja California, Baja California Sur, Sonora

Genus Stethophotopsis Pitts, 2000

367. Stethophotopsis cremauranti Pitts & Manley, 2002

Distribution Chihuahua

368. Stethophotopsis maculata Pitts, 2000

Distribution: Sonora

Subfamily Myrmosinae Fox, 1894

Tribe Kudakrumiini Krombein, 1979

Genus Myrmosula Bradley, 1917

369. Myrmosula latericarinata Wasbauer, 1973

Distribution: Baja California Sur

Species of Mutillidae incertae sedis from Mexico (Nonv2eiller, 1990; Pagliano et al., 2020)

amphissa Cameron, 1894: 325 (Sphaerophthalma); Temax in North Yucatan

antiguensis Fabricius, 1775: 396 (Mutilla)

aretes Cameron, 1896: 386 (Sphaerophthalma) Northern Sonora

cleonica Cameron, 1895: 333 (Sphaerophthalma); Temax in North Yucatan

dasygastra André, 1898: 40 (Mutilla)

erecta Fox, 1894: 93 (Sphaerophthalma)

extinctor Cameron, 1894: 322 (Sphaerophthalma); Teapa in Tabasco

ictinus Cameron, 1896: 378 (Sphaerophthalma); Northern Sonora

jaliscoensis Cameron, 1895: 346 (Sphaerophthalma); Santiago Iscuintla in Jalisco

janira Cameron, 1895: 334 (Sphaerophthalma); North Yucatan

lacuna Cameron, 1896: 380 (Sphaerophthalma)

laerma Cameron, 1896: 388 (Sphaerophthalma); Northern Sonora

luxuriosa Smith, 1879: 224 (Mutilla)

lycimnia Cameron, 1894: 327 (Sphaerophthalma); Cordova

nana Smith, 1879: 226 (Mutilla); Orizaba

oeax Cameron, 1895: 375 (Sphaerophthalma); Northern Sonora

orithyia Cameron, 1894: 325 (Sphaerophthalma); Paso del Macho

otomita Cameron, 1896 (Sphaerophthalma)

phylacis Cameron, 1896: 382 (Sphaerophthalma); Northern Sonora

semirubra Gerstaecker, 1874: 312 (Mutilla)

severa Cameron, 1896 (Sphaerophthalma)

sonorensis Cameron, 1895: 335 (Sphaerophthalma), Northern Sonora

variolosa André, 1898: 50 (Mutilla).

Nomen dubium

Invreiella megacantha (Cockerell & Casad, 1894)

Received: May 06, 2025; Accepted: June 26, 2025; Published: September 25, 2025

*Corresponding author: Fernando Varela-Hernández fernando.varela@uaem.mx

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