<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>1870-3453</journal-id>
<journal-title><![CDATA[Revista mexicana de biodiversidad]]></journal-title>
<abbrev-journal-title><![CDATA[Rev. Mex. Biodiv.]]></abbrev-journal-title>
<issn>1870-3453</issn>
<publisher>
<publisher-name><![CDATA[Instituto de Biología]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S1870-34532020000100605</article-id>
<article-id pub-id-type="doi">10.22201/ib.20078706e.2020.91.3013</article-id>
<title-group>
<article-title xml:lang="es"><![CDATA[Relaciones filogenéticas del subgénero Hypodictyon (Anura: Strabomantidae: Pristimantis) con la descripción de tres especies nuevas de la región del Chocó]]></article-title>
<article-title xml:lang="en"><![CDATA[Phylogenetic relationships of the subgenus Hypodictyon (Anura: Strabomantidae: Pristimantis) with the description of three new species from the Chocó región]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Reyes-Puig]]></surname>
<given-names><![CDATA[Carolina]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
<xref ref-type="aff" rid="Aaf"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Yánez-Muñoz]]></surname>
<given-names><![CDATA[Mario H.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Ortega]]></surname>
<given-names><![CDATA[Jhael A.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Ron]]></surname>
<given-names><![CDATA[Santiago R.]]></given-names>
</name>
<xref ref-type="aff" rid="Aff"/>
</contrib>
</contrib-group>
<aff id="Af1">
<institution><![CDATA[,Instituto Nacional de Biodiversidad Unidad de Investigación ]]></institution>
<addr-line><![CDATA[Quito ]]></addr-line>
<country>Ecuador</country>
</aff>
<aff id="Af2">
<institution><![CDATA[,Universidad San Francisco de Quito  ]]></institution>
<addr-line><![CDATA[Quito ]]></addr-line>
<country>Ecuador</country>
</aff>
<aff id="Af3">
<institution><![CDATA[,Pontificia Universidad Católica del Ecuador  ]]></institution>
<addr-line><![CDATA[Quito ]]></addr-line>
<country>Ecuador</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>00</month>
<year>2020</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>00</month>
<year>2020</year>
</pub-date>
<volume>91</volume>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.org.mx/scielo.php?script=sci_arttext&amp;pid=S1870-34532020000100605&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.org.mx/scielo.php?script=sci_abstract&amp;pid=S1870-34532020000100605&amp;lng=en&amp;nrm=iso"></self-uri><self-uri xlink:href="http://www.scielo.org.mx/scielo.php?script=sci_pdf&amp;pid=S1870-34532020000100605&amp;lng=en&amp;nrm=iso"></self-uri><abstract abstract-type="short" xml:lang="es"><p><![CDATA[Resumen Describimos 3 nuevas especies de cutines Pristimantis del subgénero Hypodictyon de la bioregión del Chocó de Colombia y Ecuador. Además, presentamos una filogenia actualizada del subgénero Hypodictyon en la que reportamos por primera vez la posición filogenética de P. calcaratus, P. colomai, P. illotus, P. kuri, P. rosadoi, P. tenebrionis y P. variabilis. Se revisó el contenido de Hypodictyon y sus 2 grupos de especies. Pristimantis afrox sp. nov. se caracteriza por la presencia de un pliegue escapular en forma de x; pliegue interocular y tubérculo longitudinal en la región interorbital; dedos del pie con pronunciados rebordes cutáneos basales en los dedos II-V; distintiva coloración inguinal y superficies ocultas de los muslos rojas. Pristimantis chocoensis sp. nov. posee dedo I de la mano igual o ligeramente más corto que el II; vientre y garganta homogéneamente crema a ligeramente manchados de café, pliegue interorbital, rebordes cutáneos laterales presentes, pliegue metatarsal interno y manchas negras hacia la inserción posterior del brazo. Pristimantis cisnerosi sp. nov. presenta distintivos huesos nasales fuertemente convexos y acampanados; narinas protuberantes proyectadas anteriormente; coloración dorsal café oscura y vientre café escarlata; tubérculos cónicos sobre el párpado y tímpano reducido.]]></p></abstract>
<abstract abstract-type="short" xml:lang="en"><p><![CDATA[Abstract We describe 3 new species of frogs Pristimantis, subgenus Hypodictyon from the Chocó of Colombia and Ecuador. Additionally, we present an updated phylogeny of the subgenus Hypodictyon on which we report, for the first time, the phylogenetic position of P. calcaratus, P. colomai, P. illotus, P. kuri, P. rosadoi, P. tenebrionis, and P. variabilis. We review the species content of Hypodictyon and its 2 species groups. Pristimantis afrox sp. nov. is characterized by bearing an x -shaped scapular fold; an interocular fold and a longitudinal tubercle in the interorbital region, toes with conspicuous lateral fringes on fingers II-V; distinctive coloration on groin and hidden surfaces of thighs red. Pristimantis chocoensis sp. nov. is characterized by having finger I equal or slightly shorter than II; venter and throat homogenously cream to slightly stained with brown, and black marks towards the insertion of the arm. Pristimantis cisnerosi sp. nov. exhibits strongly convex and flared nasal bones; protruding nostrils anteriorly; dorsal coloration dark brown and venter scarlet; bearing conical tubercles on the upper eyelid and reduced tympanum.]]></p></abstract>
<kwd-group>
<kwd lng="es"><![CDATA[Bioregión]]></kwd>
<kwd lng="es"><![CDATA[Terrarana]]></kwd>
<kwd lng="es"><![CDATA[Filogenia]]></kwd>
<kwd lng="es"><![CDATA[Pristimantis afrox sp. nov.]]></kwd>
<kwd lng="es"><![CDATA[Pristimantis chocoensis sp. nov.]]></kwd>
<kwd lng="es"><![CDATA[Pristimantis cisnerosi sp. nov]]></kwd>
<kwd lng="en"><![CDATA[Bioregion]]></kwd>
<kwd lng="en"><![CDATA[Terrarana]]></kwd>
<kwd lng="en"><![CDATA[Phylogeny]]></kwd>
<kwd lng="en"><![CDATA[Pristimantis afrox sp. nov.]]></kwd>
<kwd lng="en"><![CDATA[Pristimantis chocoensis sp. nov.]]></kwd>
<kwd lng="en"><![CDATA[Pristimantis cisnerosi sp. nov]]></kwd>
</kwd-group>
</article-meta>
</front><back>
<ref-list>
<ref id="B1">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Arteaga]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Pyron]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Peñafiel]]></surname>
<given-names><![CDATA[N.]]></given-names>
</name>
<name>
<surname><![CDATA[Romero-Barreto]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
<name>
<surname><![CDATA[Culebras]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Bustamante]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Comparative phylogeography reveals cryptic diversity and repeated patterns of cladogenesis for amphibians and reptiles in Northwestern Ecuador]]></article-title>
<source><![CDATA[Plos One]]></source>
<year>2016</year>
<volume>11</volume>
</nlm-citation>
</ref>
<ref id="B2">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Barbour]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New Central American frogs]]></article-title>
<source><![CDATA[Proceedings of the New England Zoological Club. Cambridge, Massachusetts]]></source>
<year>1928</year>
<volume>10</volume>
<page-range>25-31</page-range></nlm-citation>
</ref>
<ref id="B3">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Barrio-Amorós]]></surname>
<given-names><![CDATA[C. L.]]></given-names>
</name>
<name>
<surname><![CDATA[Guayasamin]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Hedges]]></surname>
<given-names><![CDATA[S. B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new minute Andean Pristimantis (Anura: Strabomantidae) from Venezuela]]></article-title>
<source><![CDATA[Phyllomedusa]]></source>
<year>2012</year>
<volume>11</volume>
<page-range>83-93</page-range></nlm-citation>
</ref>
<ref id="B4">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Boettger]]></surname>
<given-names><![CDATA[O.]]></given-names>
</name>
</person-group>
<source><![CDATA[Katalog der Batrachier-Sammlung im Museum der Senckenbergischen Naturforschenden Gesellshaft in Frankfurt am Main]]></source>
<year>1892</year>
<publisher-loc><![CDATA[Frankfurt ]]></publisher-loc>
<publisher-name><![CDATA[Gebrüder Knauer]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B5">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Boulenger]]></surname>
<given-names><![CDATA[G. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Descriptions of new reptiles and batrachians from Colombia]]></article-title>
<source><![CDATA[Annals and Magazine of Natural History, Series 6]]></source>
<year>1896</year>
<volume>17</volume>
<page-range>16-21</page-range></nlm-citation>
</ref>
<ref id="B6">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Boulenger]]></surname>
<given-names><![CDATA[G. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[An account of the reptiles and batrachians collected by Mr. W. F. H. Rosenberg in Western Ecuador]]></article-title>
<source><![CDATA[Proceedings of the Zoological Society of London, 1898]]></source>
<year>1898</year>
<page-range>107-26</page-range></nlm-citation>
</ref>
<ref id="B7">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Boulenger]]></surname>
<given-names><![CDATA[G. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Descriptions of new batrachians and reptiles discovered by Mr. M.G. Palmer in south-western Colombia]]></article-title>
<source><![CDATA[Annals and Magazine of Natural History, Series 8]]></source>
<year>1908</year>
<volume>2</volume>
<page-range>515-22</page-range></nlm-citation>
</ref>
<ref id="B8">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Brito]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Almendáriz]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Batallas]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Ron]]></surname>
<given-names><![CDATA[S. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Nueva especie de rana bromelícola del género Pristimantis (Amphibia: Craugastoridae), meseta de la cordillera del Cóndor, Ecuador]]></article-title>
<source><![CDATA[Papeis Avulsos de Zoologia, São Paulo]]></source>
<year>2017</year>
<volume>57</volume>
<page-range>177-95</page-range></nlm-citation>
</ref>
<ref id="B9">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Brito]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
<name>
<surname><![CDATA[Pozo-Zamora]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Una nueva especie de rana terrestre del género Pristimantis (Amphibia: Craugastoridae), de la Cordillera de Kutukú, Ecuador]]></article-title>
<source><![CDATA[Papéis Avulsos de Zoologia, São Paulo]]></source>
<year>2013</year>
<volume>53</volume>
<page-range>315-25</page-range></nlm-citation>
</ref>
<ref id="B10">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Canedo]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Haddad]]></surname>
<given-names><![CDATA[C. F.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Phylogenetic relationships within anuran clade Terrarana, with emphasis on the placement of Brazilian Atlantic rainforest frogs genus Ischnocnema (Anura: Brachycephalidae)]]></article-title>
<source><![CDATA[Molecular Phylogenetics and Evolution]]></source>
<year>2012</year>
<volume>65</volume>
<page-range>610-20</page-range></nlm-citation>
</ref>
<ref id="B11">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cope]]></surname>
<given-names><![CDATA[E. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Fourth contribution to the herpetology of tropical America]]></article-title>
<source><![CDATA[Proceedings of the Academy of Natural Sciences of Philadelphia]]></source>
<year>1866</year>
<volume>18</volume>
<page-range>123-32</page-range></nlm-citation>
</ref>
<ref id="B12">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cope]]></surname>
<given-names><![CDATA[E. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[On the Batrachia and Reptilia of Costa Rica]]></article-title>
<source><![CDATA[Journal of the Academy of Natural Sciences of Philadelphia, Series 2]]></source>
<year>1875</year>
<volume>8</volume>
<page-range>93-154</page-range></nlm-citation>
</ref>
<ref id="B13">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Cope]]></surname>
<given-names><![CDATA[E. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A contribution to the herpetology of Mexico]]></article-title>
<source><![CDATA[Proceedings of the American Philosophical Society]]></source>
<year>1885</year>
<volume>22</volume>
<page-range>380-404</page-range></nlm-citation>
</ref>
<ref id="B14">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Crawford]]></surname>
<given-names><![CDATA[A. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Cruz]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Griffith]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
<name>
<surname><![CDATA[Ross]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
<name>
<surname><![CDATA[Ibáñez]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Lips]]></surname>
<given-names><![CDATA[K. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[DNA barcoding applied to ex situ tropical amphibian conservation programme reveals cryptic diversity in captive populations]]></article-title>
<source><![CDATA[Molecular Ecology Resources]]></source>
<year>2013</year>
<volume>13</volume>
<page-range>1005-18</page-range></nlm-citation>
</ref>
<ref id="B15">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Crawford]]></surname>
<given-names><![CDATA[A. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Ryan]]></surname>
<given-names><![CDATA[M. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Jaramillo]]></surname>
<given-names><![CDATA[C. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species of Pristimantis (Anura: Strabomantidae) from the Pacific coast ofthe Darien Province, Panama, with a molecular analysis of its phylogenetic position]]></article-title>
<source><![CDATA[Herpetologica]]></source>
<year>2010</year>
<volume>66</volume>
<page-range>192-206</page-range></nlm-citation>
</ref>
<ref id="B16">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Darst]]></surname>
<given-names><![CDATA[C. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Cannatella]]></surname>
<given-names><![CDATA[D. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Novel relationships among hyloid frogs inferred from 12S and 16S mitochondrial DNA sequences]]></article-title>
<source><![CDATA[Molecular Phylogenetics and Evolution]]></source>
<year>2004</year>
<volume>31</volume>
<page-range>462-75</page-range></nlm-citation>
</ref>
<ref id="B17">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Duellman]]></surname>
<given-names><![CDATA[W.]]></given-names>
</name>
<name>
<surname><![CDATA[Lehr]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
</person-group>
<source><![CDATA[Terrestrial breeding frogs (Strabomantidae) in Peru]]></source>
<year>2009</year>
<publisher-loc><![CDATA[Alemania ]]></publisher-loc>
<publisher-name><![CDATA[NTV Science]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B18">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Echeverry]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Harper]]></surname>
<given-names><![CDATA[G. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Fragmentación y deforestación como indicadores del estado de los ecosistemas en el Corredor de Conservación Choco-Manabí (Colombia-Ecuador)]]></article-title>
<source><![CDATA[Recursos Naturales y Ambiente]]></source>
<year>2009</year>
<volume>58</volume>
<page-range>78-88</page-range></nlm-citation>
</ref>
<ref id="B19">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Firkowski]]></surname>
<given-names><![CDATA[C. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Bornschein]]></surname>
<given-names><![CDATA[M. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Ribeiro]]></surname>
<given-names><![CDATA[L. F.]]></given-names>
</name>
<name>
<surname><![CDATA[Pie]]></surname>
<given-names><![CDATA[M. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Species delimitation, phylogeny and evolutionary demography of co-distributed, montane frogs in the southern Brazilian Atlantic Forest]]></article-title>
<source><![CDATA[Molecular Phylogenetics and Evolution]]></source>
<year>2016</year>
<volume>100</volume>
<page-range>345-60</page-range></nlm-citation>
</ref>
<ref id="B20">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Flores]]></surname>
<given-names><![CDATA[G.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Two new species of Ecuadorian Eleutherodactylus (Leptodactylidae) of the E. crucifer assembly]]></article-title>
<source><![CDATA[Journal of Herpetology]]></source>
<year>1988</year>
<volume>22</volume>
<page-range>34-11</page-range></nlm-citation>
</ref>
<ref id="B21">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Frost]]></surname>
<given-names><![CDATA[D. R.]]></given-names>
</name>
</person-group>
<source><![CDATA[Amphibian species of the world: an online reference. Version 6.0]]></source>
<year>2019</year>
<publisher-loc><![CDATA[New York, USA ]]></publisher-loc>
<publisher-name><![CDATA[American Museum of Natural History]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B22">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[García]]></surname>
<given-names><![CDATA[J. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Crawford]]></surname>
<given-names><![CDATA[A. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Mendoza]]></surname>
<given-names><![CDATA[Á. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Ospina]]></surname>
<given-names><![CDATA[O.]]></given-names>
</name>
<name>
<surname><![CDATA[Cardenas]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
<name>
<surname><![CDATA[Castro]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Comparative phylogeography of direct-developing frogs (Anura: Craugastoridae: Pristimantis) in the southern Andes of Colombia]]></article-title>
<source><![CDATA[Plos One]]></source>
<year>2012</year>
<volume>7</volume>
</nlm-citation>
</ref>
<ref id="B23">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[García y Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species of frog (genus Eleutherodactylus) from a cloud forest in western Colombia]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2006</year>
<volume>1171</volume>
<page-range>39-45</page-range></nlm-citation>
</ref>
<ref id="B24">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[García]]></surname>
<given-names><![CDATA[J. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Mendoza]]></surname>
<given-names><![CDATA[Á. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Ospina]]></surname>
<given-names><![CDATA[O.]]></given-names>
</name>
<name>
<surname><![CDATA[Cardenas]]></surname>
<given-names><![CDATA[H.]]></given-names>
</name>
<name>
<surname><![CDATA[Castro]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A morphometric and molecular approach to define three closely related species of frogs of the Genus Pristimantis (Anura: Craugastoridae) from the Cordillera Occidental in Colombia]]></article-title>
<source><![CDATA[Journal of Herpetology]]></source>
<year>2014</year>
<volume>48</volume>
<page-range>220-7</page-range></nlm-citation>
</ref>
<ref id="B25">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Guayasamin]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species of Eleutherodactylus (Anura: Leptodactylidae) from the northwestern lowlands of Ecuador]]></article-title>
<source><![CDATA[Herpetologica]]></source>
<year>2004</year>
<volume>60</volume>
<page-range>103-16</page-range></nlm-citation>
</ref>
<ref id="B26">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hammer]]></surname>
<given-names><![CDATA[Ø.]]></given-names>
</name>
<name>
<surname><![CDATA[Harper]]></surname>
<given-names><![CDATA[D. A. T.]]></given-names>
</name>
<name>
<surname><![CDATA[Ryan]]></surname>
<given-names><![CDATA[P. D.]]></given-names>
</name>
</person-group>
<source><![CDATA[PAST-palaeontological statistics, ver. 1.89]]></source>
<year>2008</year>
<publisher-loc><![CDATA[Noruega ]]></publisher-loc>
<publisher-name><![CDATA[Paleontological Museum, University of Oslo]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B27">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Hedges]]></surname>
<given-names><![CDATA[S. B.]]></given-names>
</name>
<name>
<surname><![CDATA[Duellman]]></surname>
<given-names><![CDATA[W. E.]]></given-names>
</name>
<name>
<surname><![CDATA[Heinicke]]></surname>
<given-names><![CDATA[M. P.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2008</year>
<volume>1737</volume>
<page-range>1-182</page-range></nlm-citation>
</ref>
<ref id="B28">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Heinicke]]></surname>
<given-names><![CDATA[M. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Duellman]]></surname>
<given-names><![CDATA[W. E.]]></given-names>
</name>
<name>
<surname><![CDATA[Hedges]]></surname>
<given-names><![CDATA[S. B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Major Caribbean and Central American frog faunas originated by ancient oceanic dispersal]]></article-title>
<source><![CDATA[Proceedings of the National Academy of Sciences of the United States of America]]></source>
<year>2007</year>
<volume>104</volume>
<page-range>10092-7</page-range></nlm-citation>
</ref>
<ref id="B29">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Heinicke]]></surname>
<given-names><![CDATA[M. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Duellman]]></surname>
<given-names><![CDATA[W. E.]]></given-names>
</name>
<name>
<surname><![CDATA[Trueb]]></surname>
<given-names><![CDATA[L.]]></given-names>
</name>
<name>
<surname><![CDATA[Means]]></surname>
<given-names><![CDATA[D. B.]]></given-names>
</name>
<name>
<surname><![CDATA[MacCulloch]]></surname>
<given-names><![CDATA[R. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Hedges]]></surname>
<given-names><![CDATA[S. B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new frog family (Anura: Terrarana) from South America and an expanded direct-eeveloping clade revealed by molecular phylogeny]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2009</year>
<volume>2211</volume>
<page-range>1-35</page-range></nlm-citation>
</ref>
<ref id="B30">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Heinicke]]></surname>
<given-names><![CDATA[M. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Lemmon]]></surname>
<given-names><![CDATA[A. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Lemmon]]></surname>
<given-names><![CDATA[E. M.]]></given-names>
</name>
<name>
<surname><![CDATA[McGrath]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Hedges]]></surname>
<given-names><![CDATA[S. B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Phylogenomic support for evolutionary relationships of New World direct-developing frogs (Anura: Terraranae)]]></article-title>
<source><![CDATA[Molecular Phylogenetics and Evolution]]></source>
<year>2017</year>
<volume>118</volume>
<page-range>145-55</page-range></nlm-citation>
</ref>
<ref id="B31">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ibáñez]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Jaramillo]]></surname>
<given-names><![CDATA[C. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Arosemena]]></surname>
<given-names><![CDATA[F. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species of Eleutherodactylus (Anura: Leptodactylidae) from Panamá]]></article-title>
<source><![CDATA[Amphibia-Reptilia]]></source>
<year>1994</year>
<volume>15</volume>
<page-range>337-41</page-range></nlm-citation>
</ref>
<ref id="B32">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Jiménez-de la Espada]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<source><![CDATA[Vertebrados del Viaje al Pacífico Verificado de 1862 a 1865 por una Comisión de Naturalistas Enviada por el Gobierno Español]]></source>
<year>1875</year>
<publisher-loc><![CDATA[Madrid ]]></publisher-loc>
<publisher-name><![CDATA[Batracios]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B33">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Katoh]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Kuma]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Toh]]></surname>
<given-names><![CDATA[H]]></given-names>
</name>
<name>
<surname><![CDATA[Miyata]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[MAFFT version 5: improvement in accuracy of multiple sequence alignment]]></article-title>
<source><![CDATA[Nucleic Acids Research]]></source>
<year>2005</year>
<volume>33</volume>
<page-range>511-8</page-range></nlm-citation>
</ref>
<ref id="B34">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Katoh]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Misawa]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Kuma]]></surname>
<given-names><![CDATA[K. I.]]></given-names>
</name>
<name>
<surname><![CDATA[Miyata]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[MAFFT: a novel method for rapid multiple sequence alignment based on fast Fourier transform]]></article-title>
<source><![CDATA[Nucleic Acids Research]]></source>
<year>2002</year>
<volume>30</volume>
<page-range>3059-66</page-range></nlm-citation>
</ref>
<ref id="B35">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Katoh]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
<name>
<surname><![CDATA[Stanley]]></surname>
<given-names><![CDATA[D. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[MAFFT multiple sequence alignment software version 7: improvements in performance and usability]]></article-title>
<source><![CDATA[Molecular Biology and Evolution]]></source>
<year>2013</year>
<volume>30</volume>
<page-range>772-80</page-range></nlm-citation>
</ref>
<ref id="B36">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lanfear]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Calcott]]></surname>
<given-names><![CDATA[B.]]></given-names>
</name>
<name>
<surname><![CDATA[Ho]]></surname>
<given-names><![CDATA[S. Y.]]></given-names>
</name>
<name>
<surname><![CDATA[Guindon]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[PartitionFinder: combined selection of partitioning schemes and substitution models for phylogenetic analyses]]></article-title>
<source><![CDATA[Molecular Biology and Evolution]]></source>
<year>2012</year>
<volume>29</volume>
<page-range>1695-701</page-range></nlm-citation>
</ref>
<ref id="B37">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lehr]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New eleutherodactyline frogs (Leptodactylidae: Pristimantis, Phrynopus) from Peru]]></article-title>
<source><![CDATA[Bulletin of the Museum of Comparative Zoology, Cambridge, Massachusetts]]></source>
<year>2007</year>
<volume>159</volume>
<page-range>145-78</page-range></nlm-citation>
</ref>
<ref id="B38">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Two new frogs of the genus Eleutherodactylus from eastern Ecuador (Amphibia: Leptodactylidae)]]></article-title>
<source><![CDATA[Journal of Herpetology]]></source>
<year>1968</year>
<volume>2</volume>
<page-range>129-35</page-range></nlm-citation>
</ref>
<ref id="B39">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The identity of the frog Eleutherodactylus conspicillatus (Günther), with descriptions of two related species from northwestern South America (Amphibia, Leptodactylidae)]]></article-title>
<source><![CDATA[Contributions in Science. Natural History Museum of Los Angeles County]]></source>
<year>1975</year>
<volume>272</volume>
<page-range>1-19</page-range></nlm-citation>
</ref>
<ref id="B40">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New species of frogs (Leptodactylidae: Eleutherodactylus) from the Pacific Versant of Ecuador]]></article-title>
<source><![CDATA[Occasional Papers of the Museum of Natural History, The University of Kansas]]></source>
<year>1976</year>
<volume>55</volume>
<page-range>1-33</page-range><publisher-loc><![CDATA[Lawrence, Kansas ]]></publisher-loc>
</nlm-citation>
</ref>
<ref id="B41">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new frog species of the Eleutherodactylus fitzingeri group from the Pacific Andean versant in Ecuador]]></article-title>
<source><![CDATA[Herpetologica]]></source>
<year>1979</year>
<volume>35</volume>
<page-range>228-33</page-range></nlm-citation>
</ref>
<ref id="B42">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species offrog (genus Eleutherodactylus: Leptodactylidae) from a cloud forest in Departamento de Santander, Colombia]]></article-title>
<source><![CDATA[Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales]]></source>
<year>1994</year>
<volume>19</volume>
<page-range>205-8</page-range></nlm-citation>
</ref>
<ref id="B43">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New species of Eleutherodactylus from the Cordillera Occidental of western Colombia with a synopsis of the distributions of species in western Colombia]]></article-title>
<source><![CDATA[Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales]]></source>
<year>1998</year>
<volume>22</volume>
<page-range>117-48</page-range></nlm-citation>
</ref>
<ref id="B44">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Lista anotada y clave para las ranas (género Eleutherodactylus) chocoanas del Valle del Cauca, y apuntes sobre las especies de la Cordillera Occidental adyacente]]></article-title>
<source><![CDATA[Caldasia]]></source>
<year>1999</year>
<volume>21</volume>
<page-range>184-202</page-range></nlm-citation>
</ref>
<ref id="B45">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New species of frogs (Eleutherodactylus: Leptodactylidae) from the Cordillera Oriental of Norte de Santander and Santander, Colombia]]></article-title>
<source><![CDATA[Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales]]></source>
<year>2003</year>
<volume>27</volume>
<page-range>449-60</page-range></nlm-citation>
</ref>
<ref id="B46">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Ardila-Robayo]]></surname>
<given-names><![CDATA[M. C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The Eleutherodactylus of the taeniatus complex in western Colombia: taxonomy and distribution]]></article-title>
<source><![CDATA[Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales]]></source>
<year>1999</year>
<volume>23</volume>
<page-range>615-24</page-range></nlm-citation>
</ref>
<ref id="B47">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Burrowes]]></surname>
<given-names><![CDATA[P. A.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The frogs of the genus Eleutherodactylus (family Leptodactylidae) at the La Planada Reserve in southwestern Colombia with descriptions of eight new species]]></article-title>
<source><![CDATA[Occasional Papers of the Museum of Natural History, University of Kansas]]></source>
<year>1990</year>
<volume>136</volume>
<page-range>1-31</page-range></nlm-citation>
</ref>
<ref id="B48">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Duellman]]></surname>
<given-names><![CDATA[W. E.]]></given-names>
</name>
</person-group>
<source><![CDATA[Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador: systematics, ecology, and biogeography]]></source>
<year>1997</year>
<publisher-name><![CDATA[University of Kansas, Museum of Natural History]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B49">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Duellman]]></surname>
<given-names><![CDATA[W. E.]]></given-names>
</name>
</person-group>
<source><![CDATA[The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes (Anura: Leptodactylidae)]]></source>
<year>1980</year>
<numero>69</numero>
<issue>69</issue>
<page-range>1-86</page-range><publisher-name><![CDATA[University of Kansas]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B50">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Miyata]]></surname>
<given-names><![CDATA[K.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Two new species of Eleutherodactylus (Amphibia: Leptodactylidae) from the lowlands and lower cloud forests of western Ecuador]]></article-title>
<source><![CDATA[Breviora]]></source>
<year>1980</year>
<volume>457</volume>
<page-range>1-12</page-range><publisher-loc><![CDATA[Cambridge, Massachusetts ]]></publisher-loc>
<publisher-name><![CDATA[Museum of Comparative Zoology]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B51">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Ruiz-Carranza]]></surname>
<given-names><![CDATA[P. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Miyamoto]]></surname>
<given-names><![CDATA[J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The identities of the colombian frogs confused with Eleutherodactylus latidiscus (Boulenger) (Amphibia: Anura, Leptodactylidae)]]></article-title>
<source><![CDATA[Occasional Papers of the Museum of Natural History, The University of Kansas]]></source>
<year>1994</year>
<numero>170</numero>
<issue>170</issue>
<page-range>1-12</page-range><publisher-loc><![CDATA[Lawrence, Kansas ]]></publisher-loc>
</nlm-citation>
</ref>
<ref id="B52">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
<name>
<surname><![CDATA[Suárez-Mayorga]]></surname>
<given-names><![CDATA[Á. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new frog (Eleutherodactylus: Leptodactylidae) from the southern part of the Cordillera Oriental of Colombia]]></article-title>
<source><![CDATA[Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales]]></source>
<year>2000</year>
<volume>24</volume>
<page-range>289-93</page-range></nlm-citation>
</ref>
<ref id="B53">
<nlm-citation citation-type="">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Maddison]]></surname>
<given-names><![CDATA[W. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Maddison]]></surname>
<given-names><![CDATA[D. R.]]></given-names>
</name>
</person-group>
<source><![CDATA[Mesquite: a modular system for evolutionary analysis. Version 2.72]]></source>
<year>2009</year>
</nlm-citation>
</ref>
<ref id="B54">
<nlm-citation citation-type="book">
<collab>MAE</collab>
<source><![CDATA[Sistema de clasificación de los ecosistemas del Ecuador continental]]></source>
<year>2013</year>
<publisher-loc><![CDATA[Quito ]]></publisher-loc>
<publisher-name><![CDATA[Subsecretaría de Patrimonio Natural]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B55">
<nlm-citation citation-type="book">
<collab>MECN</collab>
<source><![CDATA[Serie Herpetofauna del Ecuador: El Chocó Esmeraldeño]]></source>
<year>2010</year>
<publisher-loc><![CDATA[Quito ]]></publisher-loc>
<publisher-name><![CDATA[Museo Ecuatoriano de Ciencias Naturales]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B56">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Monteiro]]></surname>
<given-names><![CDATA[J. P. D. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Condez]]></surname>
<given-names><![CDATA[T. H.]]></given-names>
</name>
<name>
<surname><![CDATA[García]]></surname>
<given-names><![CDATA[P. C. D. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Comitti]]></surname>
<given-names><![CDATA[E. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Amaral]]></surname>
<given-names><![CDATA[I. B.]]></given-names>
</name>
<name>
<surname><![CDATA[Haddad]]></surname>
<given-names><![CDATA[C. F. B.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new species of Brachycephalus (Anura, Brachycephalidae) from the coast of Santa Catarina State, southern Atlantic Forest, Brazil]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2018</year>
<volume>4407</volume>
<page-range>483-505</page-range></nlm-citation>
</ref>
<ref id="B57">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Morales-Mite]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Yánez-Muñoz]]></surname>
<given-names><![CDATA[M. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Meza-Ramos]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
<name>
<surname><![CDATA[Reyes-Puig]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Herpetofauna en las reservas de la Fundación Jocotoco: Reserva Biológica Canandé]]></article-title>
<collab>MECN</collab>
<source><![CDATA[Herpetofauna en Areas Prioritarias para la Conservación: el sistema de reservas Jocotoco y Ecominga]]></source>
<year>2013</year>
<page-range>1-408</page-range><publisher-loc><![CDATA[Quito ]]></publisher-loc>
<publisher-name><![CDATA[Fundación para la Conservación Jocotoco]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B58">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Nguyen]]></surname>
<given-names><![CDATA[L. T.]]></given-names>
</name>
<name>
<surname><![CDATA[Schmidt]]></surname>
<given-names><![CDATA[H. A.]]></given-names>
</name>
<name>
<surname><![CDATA[von Haeseler]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Minh]]></surname>
<given-names><![CDATA[B. Q.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[IQ-TREE: a fast and effective stochastic algorithm for estimating maximum-likelihood phylogenies]]></article-title>
<source><![CDATA[Molecular Biology and Evolution]]></source>
<year>2014</year>
<volume>32</volume>
<page-range>268-75</page-range></nlm-citation>
</ref>
<ref id="B59">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ortega-Andrade]]></surname>
<given-names><![CDATA[H. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Venegas]]></surname>
<given-names><![CDATA[P. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new synonym for Pristimantis luscombei (Duellman and Mendelson 1995) and the description of a new species of Pristimantis from the upper Amazon basin (Amphibia: Craugastoridae)]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2014</year>
<volume>3895</volume>
<page-range>31-57</page-range></nlm-citation>
</ref>
<ref id="B60">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Padial]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Chaparro]]></surname>
<given-names><![CDATA[J. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Castroviejo-Fisher]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Guayasamin]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Lehr]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
<name>
<surname><![CDATA[Delgado]]></surname>
<given-names><![CDATA[A. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A revision of species diversity in the Neotropical genus Oreobates (Anura: Strabomantidae), with the description of three new species from the Amazonian slopes of the Andes]]></article-title>
<source><![CDATA[American Museum Novitates]]></source>
<year>2012</year>
<volume>3752</volume>
<page-range>1-55</page-range></nlm-citation>
</ref>
<ref id="B61">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Padial]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
<name>
<surname><![CDATA[Grant]]></surname>
<given-names><![CDATA[T.]]></given-names>
</name>
<name>
<surname><![CDATA[Frost]]></surname>
<given-names><![CDATA[D. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Molecular systematics of terraranas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria]]></article-title>
<source><![CDATA[Zootaxa]]></source>
<year>2014</year>
<volume>3825</volume>
<page-range>1-132</page-range></nlm-citation>
</ref>
<ref id="B62">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Páez]]></surname>
<given-names><![CDATA[N. B.]]></given-names>
</name>
<name>
<surname><![CDATA[Ron]]></surname>
<given-names><![CDATA[S. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Systematics of Huicundomantis, a new subgenus of Pristimantis (Anura, Strabomantidae) with extraordinary cryptic diversity and eleven new species]]></article-title>
<source><![CDATA[Zookeys]]></source>
<year>2019</year>
<volume>868</volume>
<page-range>1-112</page-range></nlm-citation>
</ref>
<ref id="B63">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Paz]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Ibáñez]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Lips]]></surname>
<given-names><![CDATA[K. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Crawford]]></surname>
<given-names><![CDATA[A. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Testing the role of ecology and life history in structuring genetic variation across a landscape: a trait-based phylogeographic approach]]></article-title>
<source><![CDATA[Molecular Ecology]]></source>
<year>2015</year>
<volume>24</volume>
<page-range>3723-37</page-range></nlm-citation>
</ref>
<ref id="B64">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Peters]]></surname>
<given-names><![CDATA[W. C. H.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Über eine neue Schildrötenart, Cinosternon Effeldtii und einige andere neue oder weniger bekannte Amphibien]]></article-title>
<source><![CDATA[Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlink]]></source>
<year>1873</year>
<volume>1873</volume>
<page-range>603-18</page-range></nlm-citation>
</ref>
<ref id="B65">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Pinto-Sánchez]]></surname>
<given-names><![CDATA[N. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Ibáñez]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
<name>
<surname><![CDATA[Madriñán]]></surname>
<given-names><![CDATA[S.]]></given-names>
</name>
<name>
<surname><![CDATA[Sanjur]]></surname>
<given-names><![CDATA[O. I.]]></given-names>
</name>
<name>
<surname><![CDATA[Bermingham]]></surname>
<given-names><![CDATA[E.]]></given-names>
</name>
<name>
<surname><![CDATA[Crawford]]></surname>
<given-names><![CDATA[A. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[The Great American Biotic Interchange in frogs: multiple and early colonization of Central America by the South American genus Pristimantis (Anura: Craugastoridae)]]></article-title>
<source><![CDATA[Molecular Phylogenetics and Evolution]]></source>
<year>2012</year>
<volume>62</volume>
<page-range>954-72</page-range></nlm-citation>
</ref>
<ref id="B66">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Pyron]]></surname>
<given-names><![CDATA[R. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Wiens]]></surname>
<given-names><![CDATA[J. J.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians]]></article-title>
<source><![CDATA[Molecular Phylogenetics and Evolution]]></source>
<year>2011</year>
<volume>61</volume>
<page-range>543-83</page-range></nlm-citation>
</ref>
<ref id="B67">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ron]]></surname>
<given-names><![CDATA[S. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Merino-Viteri]]></surname>
<given-names><![CDATA[A.]]></given-names>
</name>
<name>
<surname><![CDATA[Ortiz]]></surname>
<given-names><![CDATA[D. A.]]></given-names>
</name>
</person-group>
<source><![CDATA[Anfibios del Ecuador]]></source>
<year>2019</year>
<publisher-name><![CDATA[Museo de Zoología]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B68">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Rueda-Almonacid]]></surname>
<given-names><![CDATA[J. V.]]></given-names>
</name>
<name>
<surname><![CDATA[Lynch]]></surname>
<given-names><![CDATA[J. D.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Una nueva especie de Eleutherodactylus (Amphibia: Leptodactylidae) para la Cordillera Oriental de Colombia]]></article-title>
<source><![CDATA[Lozania]]></source>
<year>1983</year>
<volume>42</volume>
<page-range>1-6</page-range></nlm-citation>
</ref>
<ref id="B69">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Ryan]]></surname>
<given-names><![CDATA[M. J.]]></given-names>
</name>
<name>
<surname><![CDATA[Lips]]></surname>
<given-names><![CDATA[K. R.]]></given-names>
</name>
<name>
<surname><![CDATA[Giermakowski]]></surname>
<given-names><![CDATA[J. T.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[New species of Pristimantis (Anura: Terrarana: Strabomantinae) from Lower Central America]]></article-title>
<source><![CDATA[Journal of Herpetology]]></source>
<year>2010</year>
<volume>44</volume>
<page-range>193-200</page-range></nlm-citation>
</ref>
<ref id="B70">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Savage]]></surname>
<given-names><![CDATA[J. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new bromeliad frog of the genus Eleutherodactylus from Costa Rica]]></article-title>
<source><![CDATA[Bulletin of the Southern California Academy of Sciences]]></source>
<year>1965</year>
<volume>64</volume>
<page-range>106-10</page-range></nlm-citation>
</ref>
<ref id="B71">
<nlm-citation citation-type="">
<collab>SIAC</collab>
<source><![CDATA[Sistema de Información ambiental de Colombia]]></source>
<year>2019</year>
</nlm-citation>
</ref>
<ref id="B72">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Sierra]]></surname>
<given-names><![CDATA[R.]]></given-names>
</name>
</person-group>
<source><![CDATA[Patrones y factores de deforestación en el Ecuador continental, 1990-2010. Y un acercamiento a los próximos 10 años]]></source>
<year>2013</year>
<publisher-loc><![CDATA[Quito ]]></publisher-loc>
<publisher-name><![CDATA[Conservación Internacional Ecuador]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B73">
<nlm-citation citation-type="">
<collab>SPSS Inc</collab>
<source><![CDATA[SPSS statistics for Windows, version 17.0]]></source>
<year>2008</year>
<publisher-loc><![CDATA[Chicago ]]></publisher-loc>
</nlm-citation>
</ref>
<ref id="B74">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Waddell]]></surname>
<given-names><![CDATA[E. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Crotti]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Lougheed]]></surname>
<given-names><![CDATA[S. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Cannatella]]></surname>
<given-names><![CDATA[D. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Elmer]]></surname>
<given-names><![CDATA[K. R.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Hierarchies of evolutionary radiation in the world's most species rich vertebrate group, the Neotropical Pristimantis leaf litter frogs]]></article-title>
<source><![CDATA[Systematics and Biodiversity]]></source>
<year>2018</year>
<volume>16</volume>
<page-range>807-19</page-range></nlm-citation>
</ref>
<ref id="B75">
<nlm-citation citation-type="book">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Yánez-Muñoz]]></surname>
<given-names><![CDATA[M.]]></given-names>
</name>
<name>
<surname><![CDATA[Meza-Ramos]]></surname>
<given-names><![CDATA[P.]]></given-names>
</name>
<name>
<surname><![CDATA[Cisneros-Heredia]]></surname>
<given-names><![CDATA[D.]]></given-names>
</name>
<name>
<surname><![CDATA[Ortega-Andrade]]></surname>
<given-names><![CDATA[H. M.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Claves de identificación y lista de chequeo de la herpetofauna del Chocó esmeraldeño]]></article-title>
<collab>MECN</collab>
<source><![CDATA[Serie Herpetofauna del Ecuador: El Chocó Esmeraldeño]]></source>
<year>2010</year>
<page-range>57-94</page-range><publisher-loc><![CDATA[Quito ]]></publisher-loc>
<publisher-name><![CDATA[Museo Ecuatoriano de Ciencias Naturales]]></publisher-name>
</nlm-citation>
</ref>
<ref id="B76">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Yánez-Muñoz]]></surname>
<given-names><![CDATA[M. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Reyes-Puig]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
<name>
<surname><![CDATA[Reyes-Puig]]></surname>
<given-names><![CDATA[J. P.]]></given-names>
</name>
<name>
<surname><![CDATA[Velasco]]></surname>
<given-names><![CDATA[J. A.]]></given-names>
</name>
<name>
<surname><![CDATA[Ayala-Varela]]></surname>
<given-names><![CDATA[F.]]></given-names>
</name>
<name>
<surname><![CDATA[Torres-Carvajal]]></surname>
<given-names><![CDATA[O.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[A new cryptic species of Anolis lizard from northwestern South America (Iguanidae, Dactyloinae)]]></article-title>
<source><![CDATA[Zookeys]]></source>
<year>2018</year>
<volume>794</volume>
<page-range>135</page-range></nlm-citation>
</ref>
<ref id="B77">
<nlm-citation citation-type="journal">
<person-group person-group-type="author">
<name>
<surname><![CDATA[Yánez-Muñoz]]></surname>
<given-names><![CDATA[M. H.]]></given-names>
</name>
<name>
<surname><![CDATA[Sánchez-Nivicela]]></surname>
<given-names><![CDATA[J. C.]]></given-names>
</name>
<name>
<surname><![CDATA[Reyes-Puig]]></surname>
<given-names><![CDATA[C.]]></given-names>
</name>
</person-group>
<article-title xml:lang=""><![CDATA[Tres nuevas especies de ranas terrestres Pristimantis (Anura: Craugastoridae) de la Provincia de El Oro, Ecuador]]></article-title>
<source><![CDATA[Avances en Ciencias e Ingeníerias, Sección B]]></source>
<year>2016</year>
<volume>8</volume>
<page-range>5-25</page-range></nlm-citation>
</ref>
</ref-list>
</back>
</article>
